WMSDB -Worldwide Mollusc Species Data Base - Family CLAVATULIDAE

WMSDB - Worldwide Mollusc Species Data Base

Family: CLAVATULIDAE Author: Claudio Galli - [email protected]

(updated 03/mag/2015)

Class: GASTROPODA --- Clade: CAENOGASTROPODA-HYPSOGASTROPODA-NEOGASTROPODA-CONOIDEA ------ Family: CLAVATULIDAE Gray, 1853 (Sea) - Alphabetic order - when first name is in bold the species has images Taxa=321, Genus=16, Subgenus=4, Species=136, Subspecies=25, Synonyms=139, Images=104

aculeiformis, Perrona aculeiformis (J.B.P.A. Lamarck, 1816) aculeiformis, Pusionella aculeiformis J.B.P.A. Lamarck, 1816 - syn of: Perrona aculeiformis (J.B.P.A. Lamarck, 1816) aculeiformis intuslirata, Perrona aculeiformis intuslirata H. Strebel, 1914 acuticarinata, Makiyamaia subdeclivis acuticarinata T. Shuto, 1961 aethiopica, Turricula aethiopica (K.H.J. Thiele, 1925) aglaophanes, Clionella aglaophanes (R.B. Watson, 1882) agulhasensis, Clionella agulhasensis K.H.J. Thiele, 1925 - syn of: Clionella vilma (K.H.J. Thiele, 1925) ahuiri, Clavatula ahuiri T. Cossignani & R. Ardovini, 2014 albocincta, Pusionella albocincta (S.A.A. Petit De La Saussaye, 1851) albocincta, Clionella halistrepta albocincta W.H. Turton, 1932 amancicoi, Scaevatula amancicoi E. Rolán & C. Fernández, 1992 - syn of: Scaevatula amancioi E. Rolán & F. Fernandes, 1993 amancioi, Scaevatula amancioi E. Rolán & F. Fernandes, 1993 amianta, Fusiturris amianta (P. Dautzenberg, 1912) amicta, Surcula amicta E.A. Smith, 1877 - syn of: Turris amicta (E.A. Smith, 1877) amplisulcus, Turricula amplisulcus (K.H. Barnard, 1958) angulosa, Pusionella nifat angulosa H. Strebel, 1914 annielonae, Clavatula annielonae F. Nolf & J. Verstraeten, 2007 - syn of: Drillia annielonae F. Nolf & J. Verstraeten, 2007 asamusiensis, Clavatula asamusiensis S. Nomura & N. Zinbo, 1940 assimilans, Clionella assimilans W.H. Turton, 1932 - syn of: Clavatula tripartita (H.C. Weinkauff, 1876) atjehensis, Turricula tornata atjehensis C.H. Oostingh, 1938 australis, Surcula australis A.F.P.M. de Roissy, 1805 - syn of: Nihonia australis (A.F.P.M. de Roissy, 1805) bairstowi, Clionella bairstowi G.B. III Sowerby, 1886 - syn of: Psittacodrillia bairstowi (G.B. III Sowerby, 1886) balteata, Fusiturris balteata L.C. Kiener, 1840 - syn of: Fusiturris undatiruga (Ant. Bivona-Bernardi, 1838) biconica, Surcula biconica M.M. Schepman, 1913 - syn of: Anticlinura biconica (M.M. Schepman, 1913) bijubata, Turricula bijubata L.A. Reeve, 1843 - syn of: Turridrupa bijubata (L.A. Reeve, 1843) bimarginata, Clavatula bimarginata (J.B.P.A. Lamarck, 1822) boletus, Clionella boletus P.F. Röding, 1798 - syn of: Clionella sinuata (I. von Born, 1778) bornii, Clionella sinuata bornii (E.A. Smith, 1877) bornii, Clionella bornii E.A. Smith, 1877 - syn of: Clionella sinuata bornii (E.A. Smith, 1877) bouvieri, Surcula bouvieri F.P. Jousseaume, 1898 brachytoma, Surcula brachytoma M.M. Schepman, 1913 - syn of: Belomitra brachytoma (M.M. Schepman, 1913) buccinatus, Pusionella buccinatus J.B.P.A. Lamarck, 1822 - syn of: Pusionella vulpina (I. von Born, 1780) caerulea, Clavatula caerulea caffra, Clionella caffra E.A. Smith, 1882 - syn of: Drillia caffra (E.A. Smith, 1882) candida, Pusionella valida candida (R.A. Philippi, 1848) carpenteriana, Surcula carpenteriana W.M. Gabb, 1865 carpenteriana tryoniana, Surcula carpenteriana tryoniana W.M. Gabb, 1866 casta, Turricula casta H. Adams, 1872 - syn of: Vexillum castum (H. Adams, 1872) catelini, Pusionella catelini S.A.A. Petit De La Saussaye, 1851 - syn of: Perrona aculeiformis (J.B.P.A. Lamarck, 1816) catena, Turricula catena (L.A. Reeve, 1843) ceylonica, Turricula ceylonica (E.A. Smith, 1877) christianae, Clavatula christianae F. Nolf, 2011 cincta, Surcula cincta J.B.P.A. Lamarck, 1822 - syn of: Turridrupa cincta (J.B.P.A. Lamarck, 1822) clandestina, Gemmuloborsonia clandestina N. Puillandre, Cruaud & Y.I. Kantor, 2010 coelata, Clavatula coelata R.B. Hinds, 1843 - syn of: Crassispira coelata (R.B. Hinds, 1843) colini, Clavatula colini H. von Maltzan, 1883 colorata, Gemmuloborsonia colorata (A.V. Sysoev & P. Bouchet, 2001) compacta, Pusionella compacta H. Strebel, 1914 confusa, Clionella confusa E.A. Smith, 1906 congoensis, Clavatula congoensis F. Nolf & J. Verstraeten, 2008 conica, Clavatula muricata conica (F.E. Edwards, 1857) conica, Clavatula conica F.E. Edwards, 1857 - syn of: Clavatula muricata conica (F.E. Edwards, 1857) conspicienda, Clionella conspicienda É.A.A. Locard, 1897 - syn of: Belomitra quadruplex (R.B. Watson, 1882) contorta, Turricula contorta G. Perry, 1811 - syn of: Turricula javana (C. Linnaeus, 1767) coreanica, Makiyamaia coreanica (A. Adams & L.A. Reeve, 1850) cornulabrum, Makiyamaia cornulabrum J.T. Kuroda, 1961

coronata, Clavatula coronata J.B.P.A. Lamarck, 1801 coronatus, Clavatula coronatus O.A.L. Mörch, 1852 - syn of: Clavatula regia (P.F. Röding, 1798) corrugata, Fusiturris corrugata L.C. Kiener, 1839 - syn of: Fusiturris undatiruga (Ant. Bivona-Bernardi, 1838) cossignanii, Clavatula cossignanii R. Ardovini, 2004 costata, Clionella costata (W.J. Swainson, 1840) crebricostata, Clavatula crebricostata R.B. Hinds, 1843 crenularis, Clavatula crenularis J.B.P.A. Lamarck, 1816 - syn of: Ptychobela nodulosa (J.F. Gmelin, 1791) culleni, Turricula culleni A. Dey, 1962 debilis, Clavatula debilis (R.B. Hinds, 1843) decorata, Clavatula decorata G.B. III Sowerby, 1916 delicatulina, Clionella delicatulina É.A.A. Locard, 1897 - syn of: Belomitra quadruplex (R.B. Watson, 1882) delicatulina costulata, Clionella delicatulina costulata É.A.A. Locard, 1897 - syn of: Belomitra quadruplex (R.B. Watson, 1882) delphinae, Clavatula delphinae F. Nolf, 2008 denansi, Clavatula denansi G.B. III Sowerby, 1914 denizi, Trachydrillia denizi F. Nolf & F. Swinnen, 2010 diadema, Clavatula diadema (L.C. Kiener, 1840) didyma, Gemmuloborsonia didyma A.V. Sysoev & P. Bouchet, 1996 digueti, Turricula digueti J.F. Mabille, 1898 dolenta, Surcula dolenta W.H. Dall, 1908 - syn of: Fusiturricula dolenta (W.H. Dall, 1908) donata, Clavatula donata R.B. Hinds, 1843 duplaris, Clionella duplaris C. Hedley, 1922 - syn of: Wairarapa duplaris (C. Hedley, 1922) elizabethae, Clionella elizabethae P. Bartsch, 1915 - syn of: Nquma rousi (G.B. III Sowerby, 1886) elstoni, Toxiclionella elstoni (K.H. Barnard, 1962) erecta, Clavatula erecta W.H. Turton, 1932 ericea, Clavatula ericea R.B. Hinds, 1843 - syn of: Lioglyphostoma ericea (R.B. Hinds, 1843) exasperata, Turricula exasperata J.F. Gmelin, 1791 - syn of: Vexillum exasperatum (J.F. Gmelin, 1791) extensesculpta, Clavatula kraepelini extensesculpta (H. Strebel, 1914) faurei, Turricula faurei (K.H. Barnard, 1958) ferruginea, Fusiturris pluteata ferruginea (H. von Maltzan, 1883) filograna, Clavatula filograna N.H. Odhner, 1923 fimbriata, Clavatula fimbriata R.B. Hinds, 1843 - syn of: Daphnellopsis fimbriata (R.B. Hinds, 1843) flammea, Turricula flammea H.C.F. Schumacher, 1817 - syn of: Turricula tornata (L.W. Dillwyn, 1817) flammulata, Clavatula flammulata J. Knudsen, 1952 flavidula, Clavatula flavidula J.B.P.A. Lamarck, 1822 - syn of: Clathrodrillia flavidula (J.B.P.A. Lamarck, 1822) fulminata, Turricula tornata fulminata (L.C. Kiener, 1839) fulminata, Turricula fulminata L.C. Kiener, 1839 - syn of: Turricula tornata fulminata (L.C. Kiener, 1839) fusca, Tomellana hupferi fusca H. Strebel, 1912 gabonensis, Clavatula gabonensis J.C. Melvill, 1923 gatchensis, Surcula gatchensis R.P.J. Hervier, 1896 - syn of: Turridrupa gatchensis (R.P.J. Hervier, 1896) gemmulaeformis, Turricula gemmulaeformis (K.H.J. Thiele, 1925) ghanaensis, Pusionella ghanaensis F. Boyer & P. Ryall, 2006 gracilior, Clavatula gracilior G.B. II Sowerby, 1870 gracilis, Tomellana lineata gracilis H. Strebel, 1912 gradata, Surcula gradata K.H.J. Thiele, 1925 - syn of: Leucosyrinx verrilli W.H. Dall, 1881 granobalteus, Turricula granobalteus (C. Hedley, 1922) gravis, Makiyamaia gravis (R.B. Hinds, 1843) gravis, Clavatula gravis (R.N. Kilburn, 1985) - syn of: Makiyamaia gravis (R.B. Hinds, 1843) gravis, Clavatula gravis R.B. Hinds, 1843 - syn of: Makiyamaia gravis (R.B. Hinds, 1843) griffithii, Clavatula griffithii J.E. Gray, 1834 - syn of: Ptychobela griffithii (J.E. Gray, 1834) haasi, Pusionella haasi P. Dautzenberg, 1912 - syn of: Pusionella valida (R.W. Dunker, 1852) hadromeres, Melatoma hadromeres J.C. Melvill, 1927 - syn of: Buchema hadromeres (J.C. Melvill, 1927) halidoma, Clionella halidoma P. Bartsch, 1915 - syn of: Clionella semicostata (L.C. Kiener, 1840) haliplex, Toxiclionella haliplex (P. Bartsch, 1915) halistrepta, Clionella halistrepta (P. Bartsch, 1915) halistrepta albocincta, Clionella halistrepta albocincta W.H. Turton, 1932 hattenbergeri, Clavatula hattenbergeri F. Nolf & J. Verstraeten, 2008 helena, Clavatula helena P. Bartsch, 1915 hondoana, Turricula hondoana W.H. Dall, 1925 - syn of: Aforia circinata (W.H. Dall, 1873) horrenda, Surcula horrenda R.B. Watson, 1886 - syn of: Stenodrillia horrenda (R.B. Watson, 1886) hottentota, Clavatula hottentota E.A. Smith, 1882 - syn of: Tylotiella hottentota (E.A. Smith, 1882) hottentota, Turricula hottentota K.H.J. Thiele, 1925 - syn of: Volutomitra hottentota (K.H.J. Thiele, 1925) hupferi, Tomellana hupferi (H. Strebel, 1912) hupferi fusca, Tomellana hupferi fusca H. Strebel, 1912

impages, Toxiclionella impages (A. Adams & L.A. Reeve, 1848) imperialis, Clavatula imperialis J.B.P.A. Lamarck, 1816 impressa, Clavatula impressa R.B. Hinds, 1843 - syn of: Kylix impressa (R.B. Hinds, 1843) innotabilis, Turricula innotabilis E.A. Smith, 1890 - syn of: Vexillum innotabile (E.A. Smith, 1890) intuslirata, Perrona aculeiformis intuslirata H. Strebel, 1914 jarrigei, Gemmuloborsonia jarrigei A.V. Sysoev & P. Bouchet, 1996 javana, Turricula javana (C. Linnaeus, 1767) javana, Surcula javana C. Linnaeus, 1767 - syn of: Turricula javana (C. Linnaeus, 1767) javanus, Turricula javanus C. Linnaeus, 1767 - syn of: Turricula javana (C. Linnaeus, 1767) jenneri, Benthoclionella jenneri R.N. Kilburn, 1974 jessica, Perrona jessica J.C. Melvill, 1923 kaffraria, Clionella subventricosa kaffraria R.N. Kilburn, 1985 kaffraria, Clionella kaffraria R.N. Kilburn, 1985 - syn of: Clionella subventricosa kaffraria R.N. Kilburn, 1985 kamakurana, Turricula kamakurana H.A. Pilsbry, 1895 kamakurana ensyuensis, Turricula kamakurana ensyuensis T. Shikama & S. Hayashi, 1977 - syn of: Comitas ensyuensis (T. Shikama, 1977) karubar, Gemmuloborsonia karubar A.V. Sysoev & P. Bouchet, 1996 kelirensis, Turricula kelirensis K. Martin, 1916 knudseni, Clavatula knudseni F. Nolf & J. Verstraeten, 2007 kowiensis, Clionella rosaria kowiensis (W.H. Turton, 1932) kowiensis viridis, Clionella kowiensis viridis W.H. Turton, 1932 - syn of: Clionella rosaria (L.A. Reeve, 1846) kraepelini, Clavatula kraepelini (H. Strebel, 1914) kraepelini extensesculpta, Clavatula kraepelini extensesculpta (H. Strebel, 1914) kraepelini nana, Clavatula kraepelini nana (H. Strebel, 1914) kraepelini pallida, Clavatula kraepelini pallida H. Strebel, 1914 krausii, Clionella krausii E.A. Smith, 1877 - syn of: Clionella kraussii (E.A. Smith, 1877) kraussii, Clionella kraussii (E.A. Smith, 1877) kurodae, Makiyamaia kurodae T. Shuto & Y. Ueda, 1963 laeta, Clavatula laeta R.B. Hinds, 1843 - syn of: Splendrillia laeta (R.B. Hinds, 1843) lavinoides, Turricula lavinoides A.A. Olsson, 1922 - syn of: Fusiturricula lavinoides (A.A. Olsson, 1922) lelieuri, Clavatula lelieuri (C.A. Récluz, 1851) leschkei, Tomellana leschkei (H. Strebel, 1912) libya, Turricula libya W.H. Dall, 1919 - syn of: Tiariturris libya (W.H. Dall, 1919) liltvedi, Clionella liltvedi R.N. Kilburn, 1985 limonensis, Turricula limonensis A.A. Olsson, 1922 - syn of: Fusiturricula lavinoides (A.A. Olsson, 1922) lincta, Turricula lincta F.W. Hutton, 1885 - syn of: Microvoluta marginata (F.W. Hutton, 1885) lineata, Tomellana lineata (J.B.P.A. Lamarck, 1818) lineata gracilis, Tomellana lineata gracilis H. Strebel, 1912 lirata, Pusionella lirata A. Adams, 1853 - syn of: Clavatula milleti (S.A.A. Petit De La Saussaye, 1851) lobatopsis, Clionella lobatopsis (K.H. Barnard, 1963) lupinus, Pusionella lupinus R.A. Philippi, 1850 - syn of: Clavatula milleti (S.A.A. Petit De La Saussaye, 1851) macella, Surcula macella J.C. Melvill, 1923 - syn of: Agladrillia macella (J.C. Melvill, 1923) macilenta, Surcula macilenta J.C. Melvill, 1923 - syn of: Agladrillia macella (J.C. Melvill, 1923) major, Pusionella nifat major H. Strebel, 1914 mammillata, Makiyamaia mammillata J.T. Kuroda, 1961 marginata, Turricula marginata F.W. Hutton, 1885 - syn of: Microvoluta marginata (F.W. Hutton, 1885) marmarina, Clavatula marmarina (R.B. Watson, 1881) martensi, Clavatula martensi H.F. Von Maltzan, 1883 matthiasi, Clavatula matthiasi F. Nolf, 2008 micro, Perrona micro E. Rolán, P. Ryall & J. Horro, 2008 milleti, Clavatula milleti (S.A.A. Petit De La Saussaye, 1851) milleti subgranulatus, Clavatula milleti subgranulatus S.A.A. Petit De La Saussaye, 1851 minor, Perrona spirata minor H. Strebel, 1912 mitra, Clavatula mitra W. Wood, 1828 - syn of: Clavatula muricata (J.B.P.A. Lamarck, 1822) moosai, Gemmuloborsonia moosai A.V. Sysoev & P. Bouchet, 1996 muricata, Clavatula muricata (J.B.P.A. Lamarck, 1822) muricata conica, Clavatula muricata conica (F.E. Edwards, 1857) mystica, Clavatula mystica (L.A. Reeve, 1843) nana, Clavatula kraepelini nana (H. Strebel, 1914) nathaliae, Clavatula nathaliae F. Nolf, 2006 navarchus, Turricula navarchus (J.C. Melvill & R. Standen, 1903) nelliae, Turricula nelliae (E.A. Smith, 1877) nelliae granobalteus, Turricula nelliae granobalteus C. Hedley, 1922 - syn of: Turricula granobalteus (C. Hedley, 1922) nelliae spuria, Turricula nelliae spuria (C. Hedley, 1922)

neocaledonica, Gemmuloborsonia neocaledonica A.V. Sysoev & P. Bouchet, 1996 nereia, Clionella nereia P. Bartsch, 1915 - syn of: Clionella rosaria (L.A. Reeve, 1846) nereia, Clionella nereia W.H. Turton, 1932 - syn of: Clionella subcontracta (E.A. Smith, 1904) nifat, Pusionella nifat (J.G. Bruguière, 1789) nifat, Pusionella nifat L.C. Kiener, 1840 - syn of: Pusionella nifat angulosa H. Strebel, 1914 nifat angulosa, Pusionella nifat angulosa H. Strebel, 1914 nifat major, Pusionella nifat major H. Strebel, 1914 nifat scalarinus, Pusionella nifat scalarinus (J.B.P.A. Lamarck, 1816) nodifera, Turricula nodifera J.B.P.A. Lamarck, 1822 - syn of: Turricula javana (C. Linnaeus, 1767) notilla, Surcula notilla W.H. Dall, 1908 - syn of: Fusiturricula notilla (W.H. Dall, 1908) nux, Clionella nux L.A. Reeve, 1845 - syn of: Clionella semicostata (L.C. Kiener, 1840) obesa, Perrona obesa (L.A. Reeve, 1842) obliquicosta, Turricula obliquicosta E.C. Von Martens, 1901 - syn of: Comitas obliquicosta (E.C. Von Martens, 1901) orthopleura, Makiyamaia orthopleura (R.N. Kilburn, 1983) pallida, Clavatula kraepelini pallida H. Strebel, 1914 pardalis, Clavatula pardalis R.B. Hinds, 1843 - syn of: Anachis pardalis (R.B. Hinds, 1843) parilis, Clavatula parilis E.A. Smith, 1901 - syn of: Clavatula tripartita (H.C. Weinkauff, 1876) paupera, Turricula paupera C. Linnaeus, 1758 pellisserpentis, Scaevatula pellisserpentis S. Gofas, 1990 perksi, Surcula perksi J.C. Verco, 1896 - syn of: Epidirona perksi (J.C. Verco, 1896) perronii, Clavatula perronii (L.A. Reeve, 1843) perronii, Perrona perronii L.A. Reeve, 1843 - syn of: Clavatula perronii (L.A. Reeve, 1843) perronii reevei, Clavatula perronii reevei H. Strebel, 1912 petzyae, Clavatula petzyae F. Boyer & P. Ryall, 2006 pfefferi, Tomellana pfefferi (H. Strebel, 1912) pfefferi, Tomella pfefferi H. Strebel, 1912 - syn of: Tomellana pfefferi (H. Strebel, 1912) phallus, Clionella phallus J.F. Gmelin, 1791 - syn of: Clionella sinuata (I. von Born, 1778) pilsbryi, Turricula pilsbryi C. Hedley, 1899 - syn of: Pusia rubrum (W.J. Broderip, 1836) plicatula, Turricula plicatula W.H. Pease, 1868 - syn of: Costellaria emiliae (J.D.E. Schmeltz, 1874) pluteata, Fusiturris pluteata (L.A. Reeve, 1843) pluteata ferruginea, Fusiturris pluteata ferruginea (H. von Maltzan, 1883) polita, Turricula polita R.B. Hinds, 1843 profundorum, Turricula profundorum (E.A. Smith, 1896) proxima, Clionella proxima W.H. Turton, 1932 - syn of: Clionella rosaria (L.A. Reeve, 1846) pseudomystica, Clavatula pseudomystica F. Nolf, 2008 pulchra, Turricula pulchra A. Garrett, 1880 - syn of: Vexillum intertaeniatum (G.B. II Sowerby, 1874) punctata, Turricula punctata L.A. Reeve, 1845 - syn of: Turricula nelliae spuria (C. Hedley, 1922) pusio, Pusionella pusio I. von Born, 1778 - syn of: Pusionella nifat (J.G. Bruguière, 1789) pyramidata, Surcula pyramidata L.C. Kiener, 1840 - syn of: Drillia pyramidata (L.C. Kiener, 1840) quadruplex, Clionella quadruplex R.B. Watson, 1882 - syn of: Belomitra quadruplex (R.B. Watson, 1882) quadruplex minor, Clionella quadruplex minor É.A.A. Locard, 1897 - syn of: Belomitra quadruplex (R.B. Watson, 1882) quinteni, Clavatula quinteni F. Nolf & J. Verstraeten, 2006 rafel, Pusionella rafel P.M. Pallary, 1920 - syn of: Pusionella vulpina (I. von Born, 1780) rapulum, Pusionella rapulum G.W. Tryon, 1884 reclusianus, Pusionella reclusianus S.A.A. Petit De La Saussaye, 1851 - syn of: Pusionella valida candida (R.A. Philippi, 1848) recurvirostris, Pusionella recurvirostris F.P. Marrat, 1877 - syn of: Perrona aculeiformis (J.B.P.A. Lamarck, 1816) reevei, Clavatula perronii reevei H. Strebel, 1912 reflexa, Turricula reflexa (L.A. Reeve, 1845) regia, Clavatula regia (P.F. Röding, 1798) remorata, Pusionella remorata E.R. Sykes, 1905 retusa, Clavatula retusa R.B. Hinds, 1843 robusta, Clavatula robusta R.B. Hinds, 1843 - syn of: Cheungbeia robusta (R.B. Hinds, 1843) rosaria, Clionella rosaria (L.A. Reeve, 1846) rosaria kowiensis, Clionella rosaria kowiensis (W.H. Turton, 1932) rubida, Clavatula rubida R.B. Hinds, 1843 - syn of: Lienardia rubida (R.B. Hinds, 1843) rubiginosa, Clavatula rubiginosa R.B. Hinds, 1843 - syn of: Otitoma rubiginosa (R.B. Hinds, 1843) rubinicolor, Clionella rubinicolor L.A. Reeve, 1845 - syn of: Clionella rosaria (L.A. Reeve, 1846) rubrifasciata, Clavatula rubrifasciata (L.A. Reeve, 1845) rubrofasciata, Clavatula rubrofasciata L.A. Reeve, 1845 - syn of: Clavatula rubrifasciata (L.A. Reeve, 1845) rufanensis, Clavatula rufanensis W.H. Turton, 1932 - syn of: Clavatula taxea (P.F. Röding, 1798) sacerdos, Clavatula sacerdos (L.A. Reeve, 1845) scalaria, Makiyamaia scalaria (K.H. Barnard, 1958) scalaria, Turricula scalaria K.H. Barnard, 1958 - syn of: Makiyamaia scalaria (K.H. Barnard, 1958)

scalarinus, Pusionella nifat scalarinus (J.B.P.A. Lamarck, 1816) scalarinus, Pusionella scalarinus J.B.P.A. Lamarck, 1816 - syn of: Pusionella nifat scalarinus (J.B.P.A. Lamarck, 1816) scripta, Pusionella scripta F. Nordsieck, 1975 - syn of: Mitrella broderipi (G.B. I Sowerby, 1844) sculpta, Clavatula sculpta R.B. Hinds, 1843 - syn of: Veprecula sculpta (R.B. Hinds, 1843) semicostata, Clionella semicostata (L.C. Kiener, 1840) seminifera, Turricula seminifera A.A. Gould, 1849 - syn of: Inquisitor interrupta (J.B.P.A. Lamarck, 1816) sibogae, Makiyamaia sibogae T. Shuto, 1970 sigillata, Clionella sigillata L.A. Reeve, 1846 - syn of: Clionella semicostata (L.C. Kiener, 1840) similis, Fusiturris similis sinensis, Clavatula sinensis R.B. Hinds, 1843 - syn of: Crassispira sinensis (R.B. Hinds, 1843) sinuata, Clionella sinuata (I. von Born, 1778) sinuata bornii, Clionella sinuata bornii (E.A. Smith, 1877) smithi, Clavatula smithi J. Knudsen, 1952 solangeae, Clavatula solangeae L. Bozzetti, 2006 spicata, Clavatula spicata R.B. Hinds, 1843 - syn of: Inquisitor spicata (R.B. Hinds, 1843) spirata, Perrona spirata (J.B.P.A. Lamarck, 1816) spirata, Clavatula spirata J.B.P.A. Lamarck, 1816 - syn of: Perrona spirata (J.B.P.A. Lamarck, 1816) spirata minor, Perrona spirata minor H. Strebel, 1912 spirata weinkauffi, Perrona spirata weinkauffi H. Strebel, 1912 spuria, Turricula nelliae spuria (C. Hedley, 1922) spuria, Turricula spuria J.H.F. Link, 1807 - syn of: Turricula javana (C. Linnaeus, 1767) strebeli, Clavatula strebeli J. Knudsen, 1952 striolata, Clionella striolata W.H. Turton, 1932 subcontracta, Clionella subcontracta (E.A. Smith, 1904) subcorpulenta, Surcula subcorpulenta E.A. Smith, 1894 - syn of: Comitas symbiotes subcorpulenta (E.A. Smith, 1894) subdeclivis, Makiyamaia subdeclivis M. Yokoyama, 1926 subdeclivis acuticarinata, Makiyamaia subdeclivis acuticarinata T. Shuto, 1961 subgranulata, Pusionella subgranulata S.A.A. Petit De La Saussaye, 1851 - syn of: Clavatula milleti subgranulatus S.A.A. Petit De La Saussaye, 1851 subgranulatus, Clavatula milleti subgranulatus S.A.A. Petit De La Saussaye, 1851 subspirata, Perrona subspirata (E.C. Von Martens, 1902) subventricosa, Clionella subventricosa (E.A. Smith, 1877) subventricosa kaffraria, Clionella subventricosa kaffraria R.N. Kilburn, 1985 sulcata, Clavatula sulcata W.J. Swainson, 1840 sulcicancellata, Turricula sulcicancellata (K.H. Barnard, 1958) sumatrana, Turricula sumatrana (K.H.J. Thiele, 1925) sybaritica, Clionella sybaritica P. Bartsch, 1915 - syn of: Clionella rosaria (L.A. Reeve, 1846) taxea, Clavatula taxea (P.F. Röding, 1798) taxus, Clavatula taxus J.H. Chemnitz, 1786 - syn of: Clavatula taxea (P.F. Röding, 1798) tenerrima, Surcula tenerrima É.A.A. Locard, 1897 ex H. Fischer ms - syn of: Irenosyrinx hypomela (W.H. Dall, 1889) tessellata, Clavatula tessellata R.B. Hinds, 1843 - syn of: Kermia tessellata (R.B. Hinds, 1843) testabilis, Pusionella testabilis F.P. Jousseaume, 1896 - syn of: Daphnella rissoides (L.A. Reeve, 1843) thisbe, Surcula thisbe E.A. Smith, 1906 - syn of: Comitas thisbe (E.A. Smith, 1906) tholoides, Clionella tholoides R.B. Watson, 1882 - syn of: Kryptos tholoides (R.B. Watson, 1882) thurstoni, Turricula thurstoni (E.A. Smith, 1896) timorensis, Surcula timorensis M.M. Schepman, 1913 - syn of: Borsonia timorensis (M.M. Schepman, 1913) tornata, Turricula tornata (L.W. Dillwyn, 1817) tornata, Surcula tornata L.W. Dillwyn, 1817 - syn of: Turricula tornata (L.W. Dillwyn, 1817) tornata atjehensis, Turricula tornata atjehensis C.H. Oostingh, 1938 tornata fulminata, Turricula tornata fulminata (L.C. Kiener, 1839) tornata malmii, Turricula tornata malmii W.H. Dall, 1889 - syn of: Mioawateria malmii (W.H. Dall, 1889) torta, Fusiturris torta (P. Dautzenberg, 1912) tripartita, Clavatula tripartita (H.C. Weinkauff, 1876) tritonium, Perrona tritonium H.C.F. Schumacher, 1817 - syn of: Clavatula perronii (L.A. Reeve, 1843) tryoniana, Surcula carpenteriana tryoniana W.M. Gabb, 1866 tuberculata, Turricula tuberculata J.E. Gray, 1839 - syn of: Turricula nelliae spuria (C. Hedley, 1922) tumida, Toxiclionella tumida (G.B. III Sowerby, 1870) turriplana, Turricula turriplana (G.B. III Sowerby, 1903) turriplana, Clavatula turriplana G.B. III Sowerby, 1903 - syn of: Turricula turriplana (G.B. III Sowerby, 1903) turris, Turricula turris J.F. Gmelin, 1791 - syn of: Turricula javana (C. Linnaeus, 1767) undarituga, Fusiturris undarituga Ant. Bivona-Bernardi, 1838 - syn of: Fusiturris undatiruga (Ant. Bivona-Bernardi, 1838) undatiruga, Fusiturris undatiruga (Ant. Bivona-Bernardi, 1838) valida, Pusionella valida (R.W. Dunker, 1852) valida candida, Pusionella valida candida (R.A. Philippi, 1848)

variabilis, Surcula variabilis M.M. Schepman, 1913 - syn of: Shutonia variabilis (M.M. Schepman, 1913) vibicinus, Turricula vibicinus G.S. Helbling, 1779 - syn of: Inquisitor interrupta (J.B.P.A. Lamarck, 1816) vilma, Clionella vilma (K.H.J. Thiele, 1925) virginea, Clavatula virginea L.A. Reeve, 1843 - syn of: Clavatula muricata (J.B.P.A. Lamarck, 1822) virgineus, Clavatula virgineus (L.W. Dillwyn, 1817) vulpina, Pusionella vulpina (I. von Born, 1780) vulpinus, Pusionella vulpinus I. von Born, 1780 - syn of: Pusionella vulpina (I. von Born, 1780) wallaysi, Pusionella wallaysi S.A.A. Petit De La Saussaye, 1851 - syn of: Pusionella rapulum G.W. Tryon, 1884 wanneri, Turricula wanneri J.J. Tesch in J. Wanner, 1915 - syn of: Nassaria wanneri (J.J. Tesch in J. Wanner, 1915) weinkauffi, Perrona spirata weinkauffi H. Strebel, 1912 xanteni, Clavatula xanteni F. Nolf & J. Verstraeten, 2006

Class: GASTROPODA --- Clade: CAENOGASTROPODA-HYPSOGASTROPODA-NEOGASTROPODA-CONOIDEA ------ Family: CLAVATULIDAE Gray, 1853 Habitat= Sea, Taxonomic order Taxa=321, Genus=16, Subgenus=4, Species=136, Subspecies=25, Synonyms=139, Images=104

Genus: Benthoclionella R.N. Kilburn, 1974

(1 SP, 1 IMG)

2115504050

Benthoclionella jenneri R.N. Kilburn, 1974 Zululand, South Africa 56 mm

Genus: Clavatula J.B.P.A. Lamarck, 1801

(50 SP, 35 IMG)

5366000021

Clavatula ahuiri T. Cossignani & R. Ardovini, 2014 Morocco 15 mm

5346000075

Clavatula asamusiensis S. Nomura & N. Zinbo, 1940

No Georeferenced data available

2110054000

Clavatula bimarginata (J.B.P.A. Lamarck, 1822) Senegal

2091501250

Clavatula caerulea (H.C. Weinkauff in H.C. Weinkauff & W. Kobelt, 1875) Senegal 15 mm

synonyms: Pleurotoma caerulea H.C. Weinkauff, 1875

Drillia coerulea H.C. Weinkauff, 1875 5346000076

Clavatula christianae F. Nolf, 2011

2115504080

Clavatula colini H. von Maltzan, 1883 West Africa

No Georeferenced data available

2115513924

Clavatula congoensis F. Nolf & J. Verstraeten, 2008 Congo

2115504090

Clavatula coronata J.B.P.A. Lamarck, 1801 South Africa

2115504100

Clavatula cossignanii R. Ardovini, 2004 Senegal, Dakar 13 mm

2115513928

Clavatula crebricostata R.B. Hinds, 1843

2115504110

Clavatula debilis (R.B. Hinds, 1843) Philippines, Mactan Island 13,9 mm

synonyms: Glyphostomoides debilis R.B. Hinds, 1843 2115504120

Clavatula decorata G.B. III Sowerby, 1916 Gulf of Guinea

2115513932

Clavatula delphinae F. Nolf, 2008 Ghana

2115504130

Clavatula denansi G.B. III Sowerby, 1914 Senegal

No Georeferenced data available

2110054035

Clavatula diadema (L.C. Kiener, 1840) Gabon 25 mm

2115504140

Clavatula donata R.B. Hinds, 1843

No Georeferenced data available

2115504150

Clavatula erecta W.H. Turton, 1932

No Georeferenced data available

2115504160

Clavatula filograna N.H. Odhner, 1923 Angola 35 mm

2115504170

Clavatula flammulata J. Knudsen, 1952 Guinea Bissau 47 mm

2115504180

Clavatula gabonensis J.C. Melvill, 1923 West Africa

2115504190

Clavatula gracilior G.B. II Sowerby, 1870

No Georeferenced data available

2115513936

Clavatula hattenbergeri F. Nolf & J. Verstraeten, 2008

No Georeferenced data available

2115504200

Clavatula helena P. Bartsch, 1915 South Africa, Cape 40 mm

2110054020

Clavatula imperialis J.B.P.A. Lamarck, 1816 Angola 46,4 mm

2115513940

Clavatula knudseni F. Nolf & J. Verstraeten, 2007

No Georeferenced data available

2115504230

Clavatula kraepelini (H. Strebel, 1914) Gabon 28 mm

2115504220

Clavatula kraepelini extensesculpta (H. Strebel, 1914)

No Georeferenced data available

2115504240

Clavatula kraepelini nana

(H. Strebel, 1914)

2115504250

Clavatula kraepelini pallida

H. Strebel, 1914 Congo 18,7 mm

2110050000

Clavatula lelieuri (C.A. Récluz, 1851) Angola 23 mm

No Georeferenced data available

2115504260

Clavatula marmarina (R.B. Watson, 1881) Brasil, Pernambuco 9 mm

synonyms: Pleurotoma marmarina R.B. Watson, 1881 2115504270

Clavatula martensi H.F. Von Maltzan, 1883 West Africa

2115513944

Clavatula matthiasi F. Nolf, 2008

No Georeferenced data available

2115504610

Clavatula milleti (S.A.A. Petit De La Saussaye, 1851) Senegal 43,2 mm

synonyms: Pusionella lirata A. Adams, 1853 Fusus lupinus R.A. Philippi, 1850

Pusionella lupinus R.A. Philippi, 1850 Fusus milleti S.A.A. Petit De La Saussaye, 1851 1972650800

Clavatula milleti subgranulatus

S.A.A. Petit De La Saussaye, 1851 Senegal

synonyms: Pusionella subgranulata S.A.A. Petit De La Saussaye, 1851 Fusus subgranulatus S.A.A. Petit De La Saussaye, 1851 2110054030

Clavatula muricata (J.B.P.A. Lamarck, 1822) Angola 32 mm

synonyms: Clavatula mitra W. Wood, 1828

Clavatula virginea L.A. Reeve, 1843 2115504280

Clavatula muricata conica

(F.E. Edwards, 1857) Angola 31 mm

synonyms: Clavatula conica F.E. Edwards, 1857

2110054040

Clavatula mystica (L.A. Reeve, 1843) Western Sahara 20 mm

2115513948

Clavatula nathaliae F. Nolf, 2006

No Georeferenced data available

2115504510

Clavatula perronii (L.A. Reeve, 1843) Angola 25 mm

synonyms: Perrona perronii L.A. Reeve, 1843

Perrona tritonium H.C.F. Schumacher, 1817 2115513973

Clavatula perronii reevei

H. Strebel, 1912 Angola

2115513952

Clavatula petzyae F. Boyer & P. Ryall, 2006 Ghana

2115513956

Clavatula pseudomystica F. Nolf, 2008 Angola

2115513836

Clavatula quinteni F. Nolf & J. Verstraeten, 2006 Angola 24,5 mm

2115511060

Clavatula regia (P.F. Röding, 1798) Congo, Pointe Noire

synonyms: Clavatula coronatus O.A.L. Mörch, 1852

2115504310

Clavatula retusa R.B. Hinds, 1843

No Georeferenced data available

2115504330

Clavatula rubrifasciata (L.A. Reeve, 1845) Angola 27 mm

synonyms: Pleurotoma rubrifasciata L.A. Reeve, 1845

Clavatula rubrofasciata L.A. Reeve, 1845 2110054010

Clavatula sacerdos (L.A. Reeve, 1845) Dakar, Senegal 20 mm

synonyms: Pleurotoma sacerdos L.A. Reeve, 1845 2115504340

Clavatula smithi J. Knudsen, 1952 Angola 6 mm

2115513960

Clavatula solangeae L. Bozzetti, 2006 Lavanono, Madagascar 40 mm

synonyms: Turris solangeae L. Bozzetti, 2006 2115504350

Clavatula strebeli J. Knudsen, 1952 Angola 21,5 mm

2115504360

Clavatula sulcata W.J. Swainson, 1840

2110054015

Clavatula taxea (P.F. Röding, 1798) South Africa, Cape 78 mm

No Georeferenced data available

synonyms: Clavatula rufanensis W.H. Turton, 1932 Turris taxea P.F. Röding, 1798

Clavatula taxus J.H. Chemnitz, 1786

Murex taxus L.W. Dillwyn, 1817

Pleurotoma bipartita E.A. Smith, 1877

Clavatula parilis E.A. Smith, 1901

2110050005

Clavatula tripartita (H.C. Weinkauff, 1876) Algoa Bay, South Africa 34 mm

synonyms: Clionella assimilans W.H. Turton, 1932 Pleurotoma tripartita H.C. Weinkauff, 1876 2115504380

Clavatula virgineus (L.W. Dillwyn, 1817)

No Georeferenced data available

synonyms: Purpura virginea P.F. Röding, 1798

Murex virgineus L.W. Dillwyn, 1817 2115513964

Clavatula xanteni F. Nolf & J. Verstraeten, 2006 Angola 24,6 mm

Genus: Clionella J.E. Gray, 1847

(syn: Melatoma - 14 SP, 14 IMG) 2115513975

Clionella aglaophanes (R.B. Watson, 1882) Barbados, Saint Thomas

synonyms: Globidrillia aglaophanes R.B. Watson, 1882

Pleurotoma aglaophanes R.B. Watson, 1882 2115504400

Clionella confusa E.A. Smith, 1906 South Africa 42 mm

2115504420

Clionella costata (W.J. Swainson, 1840)

2115513974

Clionella halistrepta (P. Bartsch, 1915) South Africa 45 mm

synonyms: Crassiclava halistrepta P. Bartsch, 1915

No Georeferenced data available

2115506210

Clionella halistrepta albocincta

W.H. Turton, 1932

No Georeferenced data available

2110154010

Clionella kraussii (E.A. Smith, 1877) East London, South Africa 30 mm

synonyms: Clionella krausii E.A. Smith, 1877

Pleurotoma krausii E.A. Smith, 1877 2110154015

Clionella liltvedi R.N. Kilburn, 1985 South Africa, Cape 23,9 mm

5336000002

Clionella lobatopsis (K.H. Barnard, 1963)

No Georeferenced data available

synonyms: Ptychosyrinx lobatopsis K.H. Barnard, 1963 2110154020

Clionella rosaria (L.A. Reeve, 1846) South Africa, Jeffreys Bay 21 mm

synonyms: Clionella kowiensis viridis W.H. Turton, 1932 Pleurotoma rosaria L.A. Reeve, 1846

Clionella nereia P. Bartsch, 1915 Clionella rubinicolor L.A. Reeve, 1845

Clionella proxima W.H. Turton, 1932 Clionella sybaritica P. Bartsch, 1915

2110154050

Clionella rosaria kowiensis

(W.H. Turton, 1932)

No Georeferenced data available

Fish River mouth 11 mm

2110154025

Clionella semicostata (L.C. Kiener, 1840) South Africa, Cape Agulhas 40 mm

synonyms: Clionella halidoma P. Bartsch, 1915 Clionella sigillata L.A. Reeve, 1846

Clionella nux L.A. Reeve, 1845 Pleurotoma sigillata L.A. Reeve, 1846

Pleurotoma semicostata L.C. Kiener, 1840

2110154030

Clionella sinuata (I. von Born, 1778) South Africa, Agulhas Bank 50,9 mm

synonyms: Clionella boletus P.F. Röding, 1798 Pleurotoma buccinoides J.B.P.A. Lamarck, 1822 Buccinum sinuatum I. von Born, 1778

Terebra phallus L.A.G. d'A. Bosc, 1801

Clionella phallus J.F. Gmelin, 1791

2115504450

Clionella sinuata bornii

(E.A. Smith, 1877) South Africa, Jeffreys Bay 36,5 mm

synonyms: Pleurotoma bornii E.A. Smith, 1877

Clionella bornii E.A. Smith, 1877 2110154035

Clionella striolata W.H. Turton, 1932 South Africa, Kommetjie 20,6 mm

2115504470

Clionella subcontracta (E.A. Smith, 1904) South Africa, Jeffreys Bay 16 mm

synonyms: Clionella nereia W.H. Turton, 1932

Drillia subcontracta E.A. Smith, 1904 2110154040

Clionella subventricosa (E.A. Smith, 1877) South Africa, Cape Agulhas 22,3 mm

synonyms: Pleurotoma subventricosa E.A. Smith, 1877 2115504480

Clionella subventricosa kaffraria R.N. Kilburn, 1985 South Africa 15,9 mm

synonyms: Clionella kaffraria R.N. Kilburn, 1985 2110154045

Clionella vilma (K.H.J. Thiele, 1925) South Africa, Cape 24 mm

synonyms: Clionella agulhasensis K.H.J. Thiele, 1925

Genus: Fusiturris K.H.J. Thiele, 1929

(syn: Tyrrhenoturris - 5 SP, 5 IMG)

2115507340

Fusiturris amianta (P. Dautzenberg, 1912) Gabon 27 mm

synonyms: Pleurotoma amianta P. Dautzenberg, 1912 2114950010

Fusiturris pluteata (L.A. Reeve, 1843) Guinea Bissau 45 mm

synonyms: Pleurotoma mandarina F.P. Marrat, 1877 2115507350

Fusiturris pluteata ferruginea

(H. von Maltzan, 1883)

No Georeferenced data available

2114950000

Fusiturris similis (Ant. Bivona-Bernardi & And. BivonaBernardi, 1838) Alger Bay 53,9 mm

synonyms: Pleurotoma similis Ant. Bivona-Bernardi & And. Bivona-Bernardi, 1838 2115507360

Fusiturris torta (P. Dautzenberg, 1912) Angola 20 mm

synonyms: Pleurotoma torta P. Dautzenberg, 1912 2114950005

Fusiturris undatiruga (Ant. Bivona-Bernardi, 1838) Malaga, Spain 54 mm

synonyms: Fusiturris balteata L.C. Kiener, 1840 Fusiturris undarituga Ant. Bivona-Bernardi, 1838

Genus: Gemmuloborsonia T. Shuto, 1989

Fusiturris corrugata L.C. Kiener, 1839

(7 SP, 7 IMG)

2140012821

Gemmuloborsonia clandestina N. Puillandre, Cruaud & Y.I. Kantor, 2010 Philippines, Luzon Island 25,3 mm

Pleurotoma ondatiruga Ant. Bivona-Bernardi, 1838

2120100040

Gemmuloborsonia colorata (A.V. Sysoev & P. Bouchet, 2001) Vanuatu 61,5 mm

synonyms: Bathytoma colorata A.V. Sysoev & P. Bouchet, 2001 2115507550

Gemmuloborsonia didyma A.V. Sysoev & P. Bouchet, 1996 Indonesia, Kai Island 20,4 mm

2115507560

Gemmuloborsonia jarrigei A.V. Sysoev & P. Bouchet, 1996 Grand Passage, New Caledonia 14,4 mm

2115507570

Gemmuloborsonia karubar A.V. Sysoev & P. Bouchet, 1996 Indonesia, Tanimbar Islands 27,8 mm

2115507580

Gemmuloborsonia moosai A.V. Sysoev & P. Bouchet, 1996 Indonesia, Tanimbar Islands 32,5 mm

2115507590

Gemmuloborsonia neocaledonica A.V. Sysoev & P. Bouchet, 1996 Norfolk Ridge, New Caledonia 30,2 mm

Genus: Makiyamaia J.T. Kuroda in F.S. MacNeil, 1960

(9 SP, 5 IMG)

2110204000

Makiyamaia coreanica (A. Adams & L.A. Reeve, 1850) East China Sea 42 mm

2115508390

Makiyamaia cornulabrum J.T. Kuroda, 1961

No Georeferenced data available

2110054005

Makiyamaia gravis (R.B. Hinds, 1843) Algoa Bay, South Africa 37,1 mm

synonyms: Clavatula gravis (R.N. Kilburn, 1985) Pleurotoma gravis R.B. Hinds, 1843

Clavatula gravis R.B. Hinds, 1843 2115511640

Makiyamaia kurodae T. Shuto & Y. Ueda, 1963 Japan

2110204010

Makiyamaia mammillata J.T. Kuroda, 1961 Japan 22,1 mm

2115500055

Makiyamaia orthopleura (R.N. Kilburn, 1983) South Africa, Natal, Richards Bay 19 mm

synonyms: Turris orthopleura R.N. Kilburn, 1983 2110504075

Makiyamaia scalaria (K.H. Barnard, 1958) Indo-Pacific

synonyms: Turricula scalaria K.H. Barnard, 1958 2115511220

Makiyamaia sibogae T. Shuto, 1970 Indonesia

2110204005

Makiyamaia subdeclivis M. Yokoyama, 1926 Florida 45 mm

No Georeferenced data available

2115511650

Makiyamaia subdeclivis acuticarinata T. Shuto, 1961 Japan

Genus: Perrona H.C.F. Schumacher, 1817

(6 SP, 6 IMG)

2115504570

Perrona aculeiformis (J.B.P.A. Lamarck, 1816) Senegal 34 mm

synonyms: Pusionella aculeiformis J.B.P.A. Lamarck, 1816 Pusionella catelini S.A.A. Petit De La Saussaye, 1851

Fusus aculeiformis J.B.P.A. Lamarck, 1816 Pusionella recurvirostris F.P. Marrat, 1877

2115504560

Perrona aculeiformis intuslirata H. Strebel, 1914

No Georeferenced data available

2115504490

Perrona jessica J.C. Melvill, 1923

No Georeferenced data available

2115513968

Perrona micro E. Rolán, P. Ryall & J. Horro, 2008 Angola

2115504500

Perrona obesa (L.A. Reeve, 1842) Angola 26 mm

synonyms: Pleurotoma obesa L.A. Reeve, 1842 2115504520

Perrona spirata (J.B.P.A. Lamarck, 1816) Angola 33,7 mm

synonyms: Clavatula spirata J.B.P.A. Lamarck, 1816

Pleurotoma spirata J.B.P.A. Lamarck, 1816

2115504530

Perrona spirata minor

H. Strebel, 1912

No Georeferenced data available

2115504540

Perrona spirata weinkauffi

H. Strebel, 1912 Angola, Luanda, Cacuaco Bay 23 mm

2115504550

Perrona subspirata (E.C. Von Martens, 1902) Angola 29 mm

synonyms: Pleurotoma subspirata E.C. Von Martens, 1902

Genus: Pusionella J.E. Gray, 1847

(syn: Netrum - 8 SP, 8 IMG) 5366000020

Pusionella albocincta (S.A.A. Petit De La Saussaye, 1851) West Africa 31,2 mm

synonyms: Fusus albocinctus S.A.A. Petit De La Saussaye, 1851 2115504590

Pusionella compacta H. Strebel, 1914 Angola, Luanda, Cacuaco Bay 22 mm

2115513978

Pusionella ghanaensis F. Boyer & P. Ryall, 2006

No Georeferenced data available

2110300000

Pusionella nifat (J.G. Bruguière, 1789) Congo, Pointe Noire 49,9 mm

synonyms: Buccinum nifat J.G. Bruguière, 1789

Pusionella pusio I. von Born, 1778

2115504630

Pusionella nifat angulosa

H. Strebel, 1914

No Georeferenced data available

synonyms: Pusionella nifat L.C. Kiener, 1840 2115504640

Pusionella nifat major

H. Strebel, 1914

No Georeferenced data available

2115504650

Pusionella nifat scalarinus

(J.B.P.A. Lamarck, 1816)

No Georeferenced data available

synonyms: Pusionella scalarinus J.B.P.A. Lamarck, 1816 2115504660

Pusionella rapulum G.W. Tryon, 1884 Java, Indonesia 31,2 mm

synonyms: Pusionella wallaysi S.A.A. Petit De La Saussaye, 1851 Fusus wallaysi S.A.A. Petit De La Saussaye, 1851 2115504670

Pusionella remorata E.R. Sykes, 1905

No Georeferenced data available

2115504600

Pusionella valida (R.W. Dunker, 1852) Angola 38 mm

synonyms: Pusionella haasi P. Dautzenberg, 1912

Bullia valida R.W. Dunker, 1852 2115513820

Pusionella valida candida

(R.A. Philippi, 1848) Guinea Bissau 42 mm

synonyms: Fusus candidus R.A. Philippi, 1848

Pusionella reclusianus S.A.A. Petit De La Saussaye, 1851

2115504700

Pusionella vulpina (I. von Born, 1780) Angola 41 mm

synonyms: Fusus buccinatus J.B.P.A. Lamarck, 1822 Pusionella buccinatus J.B.P.A. Lamarck, 1822

Genus: Scaevatula S. Gofas, 1847

Pusionella rafel P.M. Pallary, 1920

(2 SP, 2 IMG) 2115504710

Scaevatula amancioi E. Rolán & F. Fernandes, 1993 Gabon, Sao Tomé Island

synonyms: Scaevatula amancicoi E. Rolán & C. Fernández, 1992 2115504720

Scaevatula pellisserpentis S. Gofas, 1990 Gabon, Sao Tomé Island 6 mm

Genus: Surcula H. Adams & A. Adams, 1853

(syn: Surgula - 2 SP, 2 IMG)

5346000165

Surcula bouvieri F.P. Jousseaume, 1898 unknown place 27,6 mm

2114300000

Surcula carpenteriana W.M. Gabb, 1865 Santa Monica Bay, California 67 mm

synonyms: Megasurcula granti P. Bartsch, 1944 2115512980

Surcula carpenteriana tryoniana W.M. Gabb, 1866 California

Genus: Tomellana W.A. Wenz, 1943

(4 SP, 3 IMG)

2128800025

Tomellana hupferi (H. Strebel, 1912) Angola, Luanda 19,37 mm

No Georeferenced data available

Pusionella vulpinus I. von Born, 1780

2128800015

Tomellana hupferi fusca

H. Strebel, 1912

No Georeferenced data available

5332000006

Tomellana leschkei (H. Strebel, 1912)

No Georeferenced data available

2110254000

Tomellana lineata (J.B.P.A. Lamarck, 1818) Angola 27 mm

synonyms: Pleurotoma lineata J.B.P.A. Lamarck, 1816 2128800035

Tomellana lineata gracilis

H. Strebel, 1912 Angola 20 mm

2128823283

Tomellana pfefferi (H. Strebel, 1912)

No Georeferenced data available

synonyms: Tomella pfefferi H. Strebel, 1912

Genus: Toxiclionella A. Powell, 1966

(syn: Toxiclonella - 4 SP, 4 IMG)

2110400015

Toxiclionella elstoni (K.H. Barnard, 1962) Transkei, South Africa 35 mm

2110400000

Toxiclionella haliplex (P. Bartsch, 1915) Transkei, South Africa 42 mm

2110400005

Toxiclionella impages (A. Adams & L.A. Reeve, 1848) South Africa 47 mm

2110400010

Toxiclionella tumida (G.B. III Sowerby, 1870) South Africa 54,5 mm

Genus: Trachydrillia F. Nolf & F. Swinnen, 2010

(1 SP, 0 IMG)

5338000020

Trachydrillia denizi F. Nolf & F. Swinnen, 2010

Genus: Turricula H.C.F. Schumacher, 1817

No Georeferenced data available

(syn: Corbulospira - 23 SP, 12 IMG)

5346000077

Turricula aethiopica (K.H.J. Thiele, 1925)

2114150000

Turricula amplisulcus (K.H. Barnard, 1958) Agulhas Bank, South Africa 29,2 mm

2115513788

Turricula catena (L.A. Reeve, 1843) Arab Emirates 29 mm

synonyms: Pleurotoma catena L.A. Reeve, 1843 2110504030

Turricula ceylonica (E.A. Smith, 1877) Sri Lanka

synonyms: Pleurotoma ceylonica E.A. Smith, 1877 2115511690

Turricula culleni A. Dey, 1962 India

No Georeferenced data available

2115510370

Turricula digueti J.F. Mabille, 1898

No Georeferenced data available

2125350045

Turricula faurei (K.H. Barnard, 1958) Algoa Bay, South Africa 21 mm

synonyms: Daphnella faurei K.H. Barnard, 1958 2115510440

Turricula gemmulaeformis (K.H.J. Thiele, 1925)

No Georeferenced data available

Indo-Pacific

2110504000

Turricula granobalteus (C. Hedley, 1922) Broome, Australia 29,5 mm

synonyms: Turricula nelliae granobalteus C. Hedley, 1922 2110504005

Turricula javana (C. Linnaeus, 1767) Congo 67 mm

synonyms: Turricula contorta G. Perry, 1811 Surcula javana C. Linnaeus, 1767 Turricula nodifera J.B.P.A. Lamarck, 1822

Pleurotoma javana J.B.P.A. Lamarck, 1816 Turricula spuria J.H.F. Link, 1807

Turricula javanus C. Linnaeus, 1767 Turricula turris J.F. Gmelin, 1791

Pleurotoma kamakurana H.A. Pilsbry, 1895

Comitas laysanica W.H. Dall, 1919

2113954005

Turricula kamakurana H.A. Pilsbry, 1895 Philippines, Masbate Island 81,6 mm

synonyms: Comitas kamakurana H.A. Pilsbry, 1895 2115511710

Turricula kelirensis K. Martin, 1916 Java

No Georeferenced data available

2110504040

Turricula navarchus (J.C. Melvill & R. Standen, 1903) Arab Emirates

synonyms: Pleurotoma navarchus J.C. Melvill & R. Standen, 1903 2110504045

Turricula nelliae (E.A. Smith, 1877) Philippines, Palawan Island 34 mm

synonyms: Pleurotoma nelliae E.A. Smith, 1877 2110504010

Turricula nelliae spuria

(C. Hedley, 1922) Taiwan 38,5 mm

synonyms: Turricula punctata L.A. Reeve, 1845 Inquisitor spurius C. Hedley, 1922

Pleurotoma tuberculata J.E. Gray, 1839 2110504060

Turricula paupera C. Linnaeus, 1758 Indonesia

synonyms: Pleurotoma paupera C. Linnaeus, 1758 2115510390

Turricula polita R.B. Hinds, 1843

No Georeferenced data available

2110504065

Turricula profundorum (E.A. Smith, 1896)

No Georeferenced data available

Indo-Pacific

2115510400

Turricula reflexa (L.A. Reeve, 1845)

No Georeferenced data available

Turricula tuberculata J.E. Gray, 1839

2110504080

Turricula sulcicancellata (K.H. Barnard, 1958)

No Georeferenced data available

2115510470

Turricula sumatrana (K.H.J. Thiele, 1925)

No Georeferenced data available

Indo-Pacific

2110504085

Turricula thurstoni (E.A. Smith, 1896)

No Georeferenced data available

Indo-Pacific

2110504090

Turricula tornata (L.W. Dillwyn, 1817) Bangladesh 97 mm

synonyms: Pleurotoma candida K.T. Menke, 1829 Surcula tornata L.W. Dillwyn, 1817

Turricula flammea H.C.F. Schumacher, 1817 Murex tornatus L.W. Dillwyn, 1817 2115510480

Turricula tornata atjehensis

C.H. Oostingh, 1938 Indonesia

2115510490

Turricula tornata fulminata

(L.C. Kiener, 1839) Oman 46 mm

synonyms: Turricula fulminata L.C. Kiener, 1839 2115510500

Turricula turriplana (G.B. III Sowerby, 1903) South Africa, Cape Agulhas 22,8 mm

synonyms: Clavatula turriplana G.B. III Sowerby, 1903

Pleurotoma turriplana G.B. III Sowerby, 1903

Family: CLAVATULIDAE

red squares show where each species has been georeferenced each square has a size of 2� an interactive version of this image can be accessed at http://www.bagniliggia.it/WMSD/FamilyMap/CLAVATULIDAEmap.htm please note that this is not the complete coverage of this family the Data Base is not exaustive and only one specimen per species has been georeferenced this map is only the graphic summary of the WMSDB Data Base up to now the total number of georeferenced species in this map is: 116 from a total of 161

Author: Claudio Galli - [email protected]