WMSDB -Worldwide Mollusc Species Data Base - Family HELICARIONIDAE

WMSDB - Worldwide Mollusc Species Data Base

Family: HELICARIONIDAE Author: Claudio Galli - [email protected]

(updated 05/mag/2015)

Class: GASTROPODA --- Clade: HETEROBRANCHIA-PULMONATA-EUPULMONATA-STYLOMMATOPHORA-SIGMURETHRA-HELICARIONOIDEA ------ Family: HELICARIONIDAE Bourguignat, 1877 (Land) - Alphabetic order - when first name is in bold the species has images Taxa=468, Genus=80, Subgenus=14, Species=311, Subspecies=20, Synonyms=42, Images=83

aberrans, Helicarion aberrans T. Iredale, 1944 achilles, Hunsteinia achilles (J.W. Brazier, 1886) acutangula, Helicarion acutangula A. Adams, 1868 acutanguloides, Parakaliella acutanguloides J.T. Kuroda & M. Azuma, 1982 adamsi, Yamatochlamys vaga adamsi J.T. Kuroda & T. Kawamoto, 1956 affinis, Geotrochus affinis (E.A. Smith, 1894) affinis, Parakaliella affinis H.A. Pilsbry & Y. Hirase, 1905 akibumii, Helicarion akibumii J.T. Kuroda, 1963 albellus, Helicarion albellus E.C. Von Martens, 1867 albocincta, Mathewsoconcha albocincta H.B. Preston, 1913 albofilosa, Sivella albofilosa (A.R.J.B. Bavay & P. Dautzenberg, 1908) alexis, Geotrochus alexis (E.A. Smith, 1895) alma, Helicarion alma T. Iredale, 1944 amiculus, Helicarion amiculus T. Iredale, 1945 ampla, Helicarion ampla O.W.H. Reinhardt, 1883 annamitica, Sesara annamitica (O.F. von Möllendorff, 1900) aquavitae, Helicarion aquavitae P.H. Scott, 1995 aquila, Helicarion aquila J.C. Cox, 1868 argentea, Harmogenanina argentea L.A. Reeve, 1852 armiti, Hunsteinia armiti (E.A. Smith, 1895) atra, Pseudaustenia atra H.H. Godwin-Austen atratus, Brevisentis atratus I. Hyman, 2007 attenuata, Laocaia attenuata A.A. Kuzminykh, 1999 austeniana, Parakaliella austeniana H.A. Pilsbry & Y. Hirase, 1901 awaensis, Helicarion awaensis H.A. Pilsbry, 1902 azumai, Helicarion azumai T. Habe, 1948 baleriana, Hemitrichiella baleriana P. Bartsch, 1939 basistriata, Trochochlamys crenulata basistriata H.A. Pilsbry & Y. Hirase, 1905 bella, Helicarion subrejectus bella H.A. Pilsbry, 1926 belli, Mathewsoconcha belli H.B. Preston, 1913 belli, Helicarion belli T. Iredale, 1944 bimaris, Parakaliella bimaris H.A. Pilsbry & Y. Hirase blainvilleana, Hemiglypta blainvilleana H.C. Lea bongaoensis, Geotrochus bongaoensis (E.A. Smith, 1894) borealis, Helicarion borealis H.A. Pilsbry, 1901 bouyei, Sesara bouyei (J.C.H. Crosse & P.H. Fischer, 1863) brazieri, Helicarion brazieri J.C. Cox, 1873 brunnea, Pseudosaphtia brunnea A.J. De Winter, 2008 brunneus, Helicarion imperator brunneus (O.F. von Möllendorff, 1901) bullaceus, Helicarion bullaceus N.H. Odhner, 1917 caelatura, Plegma caelatura (A.E.J. Férussac, 1821) caldwelli, Ctenophila caldwelli W.H. Benson, 1859 callidus, Helicarion mastersi callidus T. Iredale, 1941 caperatus, Helicarion caperatus A.A. Gould - syn of: Godwinia caperata A.A. Gould, 1864 castanea, Helicarion castanea L. Pfeiffer, 1853 castra, Sivella castra W.H. Benson, 1852 - syn of: Videna castra (W.H. Benson, 1852) catletti, Helicarion catletti J.W. Brazier, 1872 cerasina, Petalochlamys cerasina (H.A. Pilsbry, 1902) ceratodes, Helicarion ceratodes G.P.L.K. Gude, 1900 charis, Helicarion charis T. Iredale, 1944 cineraceus, Dendrotrochus cineraceus J.B. Hombron & C.H. Jacquinot, 1841 cineraceus simboana, Dendrotrochus cineraceus simboana (E.A. Smith, 1885)

circumdata, Helicarion circumdata H.A. Pilsbry, 1902 cleryi, Dendrotrochus helicinoides cleryi (C.A. Récluz, 1851) cochinchinensis, Helicarion cochinchinensis P.M.A. Morelet, 1866 - syn of: Megaustenia cochinchinensis (P.M.A. Morelet, 1866) compacta, Mathewsoconcha compacta H.B. Preston, 1913 compluviata, Orpiella compluviata (J.C. Cox, 1871) conica, Helicarion sophiae conica J.W. Brazier, 1889 conicoides, Geotrochus conicoides W. Metcalfe, 1851 conicoides parva, Geotrochus conicoides parva (E.A. Smith, 1887) conicoides subconvexa, Geotrochus conicoides subconvexa O.F. von Möllendorff connectens, Hemiglypta panayensis connectens O.F. von Möllendorff, 1898 conus, Geotrochus conus (R.A. Philippi, 1841) cookensis, Orpiella cookensis A.A. Gould coriacea, Nitor coriacea A. Manzoni, 1868 costata, Parakaliella costata H.A. Pilsbry & Y. Hirase, 1904 crassiaxis, Helicarion crassiaxis J.T. Kuroda, 1969 crenata, Helicarion crenata J.T. Kuroda & M. Azuma, 1982 crenulata, Trochochlamys crenulata G.P.L.K. Gude, 1900 crenulata basistriata, Trochochlamys crenulata basistriata H.A. Pilsbry & Y. Hirase, 1905 crenulata hotawana, Trochochlamys crenulata hotawana H.A. Pilsbry & Y. Hirase, 1905 cuvierana, Hemiglypta cuvierana (H.C. Lea, 1893) cuvieri, Helicarion cuvieri A.E.J. Férussac, 1821 cyrene, Dendrotrochus cyrene (J.C.H. Crosse, 1865) dagueneti, Nitor dagueneti L. de Folin, 1873 dahli, Conibycus dahli K.H.J. Thiele, 1928 danjoensis, Helicarion danjoensis H. Minato, 1974 decens, Japanochlamys decens H.A. Pilsbry & Y. Hirase, 1904 delecta, Helicarion delecta T. Iredale, 1944 depressa, Roybellia depressa H.B. Preston, 1913 depressa, Parakaliella depressa O.F. von Möllendorff, 1883 depressus, Helicarion depressus G.A. von Jacobi, 1898 detecta, Harmogenanina detecta (A.E.J. Férussac, 1827) diplodon, Sesara diplodon (W.H. Benson, 1859) dispositus, Helicarion dispositus T. Iredale, 1941 doenitzii, Petalochlamys doenitzii (O.W.H. Reinhardt, 1877) dulcis, Bekkochlamys dulcis H.A. Pilsbry, 1902 dulcissima, Helicarion dulcissima T. Iredale, 1944 elevata, Mathewsoconcha elevata H.B. Preston, 1913 etheridgei, Helicarion etheridgei J.W. Brazier, 1889 everetti, Geotrochus everetti (H.H. Godwin-Austen, 1891) filaris, Dendrotrochus filaris (M. Leschke, 1912) fischeri, Parmacochlea fischeri E.A. Smith, 1884 flavescens, Helicarion flavescens T. Iredale, 1944 flyensis, Hunsteinia flyensis (C. Hedley, 1894) foersteri, Hunsteinia foersteri (W. Kobelt, 1914) formosana, Petalochlamys formosana H.A. Pilsbry & Y. Hirase, 1908 - syn of: Petalochlamys formosana P.B. Schmacker & O. Böttger, 1891 formosana, Petalochlamys formosana P.B. Schmacker & O. Böttger, 1891 fouillioyi, Hemiglyptopsis fouillioyi E.J. Le Guillou, 1845 fragillima, Orpiella fragillima A.A. Gould fraterna, Trochochlamys fraterna H.A. Pilsbry, 1900 fraterna takahashii, Trochochlamys fraterna takahashii J.T. Kuroda, 1963 fraternus, Helicarion fraternus T. Iredale, 1945 fredaslini, Westracystis fredaslini G.A. Solem, 1982 freycineti, Helicarion freycineti A.E.J. Férussac, 1821 fulgens, Ovachlamys fulgens G.P.L.K. Gude, 1900 fulgida, Ovachlamys fulgida H.H. Godwin-Austen, 1907 fusaniana, Parakaliella fusaniana H.A. Pilsbry & Y. Hirase, 1909 glenrockensis, Sheaia glenrockensis I. Hyman, 2007 globosa, Hemiglypta globosa C.G. Semper, 1870 godeffroyana, Orpiella godeffroyana A. Garrett, 1872 goniozona, Trochochlamys lioconus goniozona H.A. Pilsbry & Y. Hirase, 1905 gracilior, Dendrotrochus gracilior (B. Rensch, 1935) granulosa, Saphtia granulosa A.J. De Winter, 2008 grata, Helicarion grata T. Iredale, 1944

griffithsi, Ctenophila griffithsi K.C. Emberton, 1994 guamata, Hemitrichiella guamata G.B. Sowerby gudei, Takemasaia gudei H.A. Pilsbry, 1901 habei, Parakaliella habei J.T. Kuroda, 1944 hakonensis, Helicarion hakonensis H.A. Pilsbry & Y. Hirase, 1905 hakusanus, Nipponochlamys hakusanus H.A. Pilsbry & Y. Hirase, 1907 helenae, Helicarion helenae H.H. Godwin-Austen, 1883 helenkingae, Helicarion helenkingae P.H. Scott, 1995 helicinoides, Dendrotrochus helicinoides J.B. Hombron & C.H. Jacquinot, 1841 helicinoides cleryi, Dendrotrochus helicinoides cleryi (C.A. Récluz, 1851) heraclea, Geotrochus heraclea (E.A. Smith, 1895) hercules, Hunsteinia hercules (C. Hedley, 1891) hilli, Howearion hilli (J.C. Cox, 1873) hiraseana, Helicarion hiraseana H.A. Pilsbry, 1901 hizenensis, Parakaliella hizenensis H.A. Pilsbry, 1902 hokkaidonis, Nipponochlamys hokkaidonis H.A. Pilsbry & Y. Hirase, 1905 hotawana, Trochochlamys crenulata hotawana H.A. Pilsbry & Y. Hirase, 1905 howinsulae, Epiglypta howinsulae J.C. Cox, 1873 hoyti, Irenella hoyti A. Garrett, 1872 humiliconus, Trochochlamys humiliconus H.A. Pilsbry & Y. Hirase, 1904 hyaleus, Helicarion hyaleus F.S. Bock, 1881 iloilana, Hemiglypta iloilana M. Smith, 1932 imitatrix, Helicarion imitatrix E.R. Sykes, 1900 imitratrix, Lutilodix imitratrix E.R. Sykes, 1900 imperator, Helicarion imperator (A.A. Gould, 1859) imperator brunneus, Helicarion imperator brunneus (O.F. von Möllendorff, 1901) imperfecta, Caldwellia imperfecta unknown implicata, Harmogenanina implicata G. Nevill, 1870 incerta, Parakaliella incerta A. Adams, 1868 infans, Helicarion infans (L. Pfeiffer, 1854) infredens, Sesara infredens A.A. Gould infrendens, Sesara infrendens (A.A. Gould, 1844) insculpta, Helicarion insculpta L. Pfeiffer, 1846 inversicolor, Pachystyla inversicolor A.E.J. Férussac, 1821 inversicolor mauritianella, Pachystyla inversicolor mauritianella (P.M.A. Morelet) iridis, Helicarion iridis I. Hyman, 2007 isabellensis, Geotrochus isabellensis (S.M. Souverbie, 1863) izushichitojimana, Nipponochlamys izushichitojimana H.A. Pilsbry & Y. Hirase, 1904 izushichtoensis, Trochochlamys praealta izushichtoensis H.A. Pilsbry & Y. Hirase, 1903 kagaensis, Helicarion kagaensis H.A. Pilsbry & Y. Hirase, 1902 kandai, Ovachlamys kandai M. Azuma, 1978 kermadecensis, Kermarion kermadecensis (E.A. Smith, 1873) kiiensis, Helicarion kiiensis M. Azuma, 1960 kimioi, Helicarion kimioi M. Azuma, 1960 kinabaluensis, Geotrochus kinabaluensis (E.A. Smith, 1895) kobelti, Lepidotrichia kobelti (O.F. von Möllendorff, 1890) koshikijimana, Bekkochlamys koshikijimana H.A. Pilsbry & Y. Hirase, 1904 kotosyonis, Ovachlamys kotosyonis J.T. Kuroda & T. Kano, 1941 koyamai, Helicarion koyamai J.T. Kuroda, 1953 koyanoi, Helicarion koyanoi T. Habe, 1953 kreffti, Helicarion kreffti J.C. Cox, 1864 kurodai, Helicarion kurodai T. Habe, 1957 kuroshimana, Helicarion kuroshimana Y. Hirase, 1908 labilis, Petalochlamys labilis (A.A. Gould, 1859) labillardierei, Dendrotrochus labillardierei E.A. Smith, 1884 labuanensis, Geotrochus labuanensis L. Pfeiffer, 1863 lampra, Helicarion lampra H.A. Pilsbry & Y. Hirase, 1904 lamtoensis, Saphtia lamtoensis A.J. De Winter, 2008 latior, Sivella latior (A.R.J.B. Bavay & P. Dautzenberg, 1908) layardi, Dendrotrochus layardi W.D. Hartman, 1889 leopardina, Helicarion leopardina T. Iredale, 1941 leucospira, Helicarion leucospira L. Pfeiffer, 1857 levensonia, Dupontia levensonia J. Gerlach, 2003

levis, Dupontia levis H.H. Godwin-Austen, 1908 libmanana, Hemitrichiella luteofasciata libmanana P. Bartsch, 1942 lineatus, Nipponochlamys lineatus H.A. Pilsbry & Y. Hirase, 1904 linophora, Harmogenanina linophora P.M.A. Morelet, 1860 lioconus, Trochochlamys lioconus H.A. Pilsbry & Y. Hirase, 1905 lioconus goniozona, Trochochlamys lioconus goniozona H.A. Pilsbry & Y. Hirase, 1905 lioderma, Helicarion lioderma H.A. Pilsbry, 1901 lissus, Westracystis lissus E.A. Smith, 1894 longa, Helicarion longa H.A. Pilsbry & Y. Hirase, 1905 longissima, Helicarion longissima H.A. Pilsbry & Y. Hirase, 1909 luteofasciata, Hemitrichiella luteofasciata H.C. Lea, 1840 luteofasciata libmanana, Hemitrichiella luteofasciata libmanana P. Bartsch, 1942 madagascariensis, Bathia madagascariensis G.C. Robson, 1914 maillardi, Dupontia maillardi (G.P. Deshayes, 1863) maleficus, Helicarion maleficus P. Dautzenberg, 1904 - syn of: Helicarion imperator (A.A. Gould, 1859) malleata, Nesonanina malleata B. Rensch masakii, Helicarion masakii J.T. Kuroda, 1960 mastersi, Helicarion mastersi J.C. Cox, 1868 mastersi callidus, Helicarion mastersi callidus T. Iredale, 1941 mathewsi, Helicarion mathewsi H.B. Preston, 1913 mathildae, Wilhelminaia mathildae H.B. Preston, 1913 mauritianella, Pachystyla inversicolor mauritianella (P.M.A. Morelet) mcdonaldi, Eungarion mcdonaldi J. Stanisic, 1993 medioximus, Nitor medioximus T. Iredale, 1941 megastoma, Helicarion megastoma J.C. Cox, 1868 meristotrochus, Geotrochus meristotrochus microglypta, Hemiglypta microglypta O.F. von Möllendorff, 1893 micrograpta, Bekkochlamys micrograpta H.A. Pilsbry, 1900 microstriata, Mathewsoconcha microstriata H.B. Preston, 1913 milloti, Ctenophila milloti E. Fischer-Piette, F. Blanc & F. Salvat, 1975 miranda, Helicarion miranda T. Iredale, 1944 montana, Sivella montana (O.F. von Möllendorff, 1901) monticola, Helicarion monticola J.T. Kuroda & J.M. Abe, 1980 - syn of: Helicarion monticula J.T. Kuroda & J.M. Abe, 1980 monticula, Helicarion monticula J.T. Kuroda & J.M. Abe, 1980 moolenbeeki, Rahula moolenbeeki W.J.M. Maassen, 2000 mouelti, Orpiella mouelti Kae mouelti tawale, Orpiella mouelti tawale Kae mutsuensis, Helicarion mutsuensis Y. Hirase, 1908 nachikoae, Helicarion nachikoae J.T. Kuroda, 1953 nahaensis, Parakaliella nahaensis (G.P.L.K. Gude, 1900) nevilli, Erepta nevilli A. Adams, 1867 newtoni, Ctenoglypta newtoni G. Nevill, 1871 nigra, Helicarion nigra J.R.C. Quoy & J.P. Gaimard, 1832 nikkoensis, Helicarion nikkoensis E. Sorita & M. Kawana, 1983 nissani, Owaraha nissani R.K. Dell, 1955 nitella, Dupontia nitella P.M.A. Morelet, 1868 nitidus, Petalochlamys nitidus H.A. Pilsbry & Y. Hirase, 1905 nouleti, Orpiella nouleti (E.J. Le Guillou, 1842) novaehiberniae, Nesonanina novaehiberniae J.R.C. Quoy & J.P. Gaimard, 1832 novoguineensis, Helicarion novoguineensis E.A. Smith, 1887 obesa, Laocaia obesa A.A. Kuzminykh, 1999 obscura, Geotrochus obscura obtusa, Nipponochlamys obtusa C.A. Westerlund, 1883 obtusangula, Parakaliella obtusangula O.W.H. Reinhardt, 1883 odontina, Erepta odontina P.M.A. Morelet, 1851 ogasawarana, Helicarion ogasawarana H.A. Pilsbry, 1902 okinoshimana, Trochochlamys okinoshimana H.A. Pilsbry & Y. Hirase, 1904 orthostoma, Inozonites orthostoma (L. Pfeiffer, 1845) osimakiheijii, Helicarion osimakiheijii J.T. Kuroda, 1953 otakiana, Parakaliella otakiana Y. Hirase, 1909 otareae, Irenella otareae A. Garrett pagoduloides, Parakaliella pagoduloides G.P.L.K. Gude, 1900 pallida, Geotrochus pallida (E.A. Smith, 1895)

panayensis, Hemiglypta panayensis W.J. Broderip, 1842 panayensis connectens, Hemiglypta panayensis connectens O.F. von Möllendorff, 1898 pandana, Lepidotrichia pandana (P. Bartsch, 1942) par, Petalochlamys par P.B. Schmacker & O. Böttger, 1891 paraguensis, Geotrochus paraguensis (E.A. Smith, 1893) parkini, Cucullarion parkini J. Stanisic, 1998 parva, Geotrochus conicoides parva (E.A. Smith, 1887) paviei, Sivella paviei (L.J. Morlet, 1884) perakensis, Geotrochus perakensis J.C.H. Crosse, 1879 - syn of: Ganesella acris perakensis (J.C.H. Crosse, 1879) perfragilis, Petalochlamys perfragilis (H.A. Pilsbry, 1901) perfragilis sakui, Petalochlamys perfragilis sakui H.A. Pilsbry & Y. Hirase, 1908 perlucida, Dupontia perlucida A. Adams, 1867 perrugosa, Panulena perrugosa T. Iredale, 1945 pfeifferi, Orpiella pfeifferi (R.A. Philippi, 1845) phillipii, Mathewsoconcha phillipii J.E. Gray, 1834 philyrina, Caldwellia philyrina P.M.A. Morelet, 1851 phonicus, Geotrochus phonicus J.F. Mabille, 1887 - syn of: Ganesella phonica (J.F. Mabille, 1887) phyllophila, Parakaliella phyllophila A. Adams, 1868 planilabris, Helicarion planilabris J.C. Cox, 1866 planospirus, Helicarion planospirus (L. Pfeiffer, 1853) platysoma, Roybellia platysoma E.R. Sykes, 1900 ponapensis, Dendrotrochus ponapensis H.B. Baker, 1941 ponderi, Levidens ponderi I. Hyman, 2007 porrectus, Helicarion porrectus T. Iredale, 1941 poweri, Dupontia poweri A. Adams, 1868 praealta, Trochochlamys praealta H.A. Pilsbry, 1902 praealta izushichtoensis, Trochochlamys praealta izushichtoensis H.A. Pilsbry & Y. Hirase, 1903 prensa, Helicarion prensa T. Iredale, 1944 princeps, Helicarion princeps T. Iredale, 1944 proletaria, Dupontia proletaria P.M.A. Morelet, 1860 pruinosa, Lepidotrichia pruinosa (O.F. von Möllendorff, 1890) pudibunda, Nitor pudibunda J.C. Cox, 1868 pupula, Parakaliella pupula A.A. Gould, 1859 purpurescens, Lepidotrichia purpurescens O.F. von Möllendorff pustulina, Parakaliella pustulina O.W.H. Reinhardt, 1877 quelpartensis, Helicarion quelpartensis H.A. Pilsbry & Y. Hirase, 1902 quintalae, Helicarion quintalae J.C. Cox, 1870 radiatulus, Helicarion radiatulus (O.F. von Möllendorff, 1897) rapida, Helicarion rapida J.T. Kuroda & M. Miyanaga, 1939 raricostulata, Rahula raricostulata (E.A. Smith, 1894) rejecta, Petalochlamys rejecta L. Pfeiffer, 1865 retinaculum, Helicarion retinaculum H.B. Preston, 1913 rhysa, Geotrochus rhysa S. Tillier & P. Bouchet, 1988 rimatus, Geotrochus rimatus J.J. Vermeulen, 1997 robusta, Helicarion robusta A.A. Gould, 1846 rodriguezensis, Dancea rodriguezensis J.C.H. Crosse, 1873 rosacea, Helicarion rosacea T. Iredale, 1944 rubicundus, Helicarion rubicundus Dartnall & J.A. Kershaw, 1978 rufozonata, Pachystyla rufozonata A. Adams, 1867 rugosa, Helicarion rugosa H.C. Fulton, 1910 ruida, Parakaliella ruida H.A. Pilsbry, 1901 saigonensis, Geotrochus saigonensis (J.C.H. Crosse, 1867) sakui, Petalochlamys perfragilis sakui H.A. Pilsbry & Y. Hirase, 1908 sapeca, Geotrochus sapeca P.M. Heude, 1890 - syn of: Videna sapeca (P.M. Heude, 1890) satsumana, Helicarion subcrenulata satsumana H.A. Pilsbry & Y. Hirase, 1908 satsumana, Helicarion satsumana Y. Hirase, 1908 scorpio, Orpiella scorpio A.A. Gould semicalcareus, Helicarion semicalcareus M.M. Schepman, 1896 semiglobosa, Hemiglypta semiglobosa (L. Pfeiffer) semigranosa, Hemiglypta semigranosa G.B. I Sowerby, 1832 semisculpta, Lepidotrichia semisculpta O.F. von Möllendorff semisericata, Nipponochlamys semisericata (H.A. Pilsbry, 1902) semoni, Thularion semoni E.C. Von Martens, 1894

semperi, Hemiglypta semperi O.F. von Möllendorff septentrionalis, Helicarion septentrionalis G.A. von Jacobi, 1898 serenus, Helicarion serenus H.A. Pilsbry & Y. Hirase, 1908 serica, Parakaliella serica H.A. Pilsbry, 1927 setigera, Hemitrichiella setigera G.B. Sowerby, 1898 setiliris, Ctenophila setiliris W.H. Benson, 1851 shikokuensis, Bekkochlamys shikokuensis H.A. Pilsbry & Y. Hirase, 1903 shikokuensis, Parakaliella shikokuensis J.T. Kuroda & J.M. Abe, 1980 shinanoensis, Helicarion shinanoensis H.A. Pilsbry & Y. Hirase, 1904 shiroyamaensis, Helicarion shiroyamaensis J.T. Kuroda & T. Habe, 1954 sieversi, Helicarion sieversi A. Mousson, 1876 - syn of: Phenacolimax annularis (S.E. Studer, 1820) simboana, Dendrotrochus cineraceus simboana (E.A. Smith, 1885) smithi, Parmacochlea smithi H.R. Simroth, 1898 sogdianus, Helicarion sogdianus E.C. Von Martens, 1871 - syn of: Macrochlamys sogdiana (E.C. Von Martens, 1871) solidiuscula, Orpiella solidiuscula (E.A. Smith, 1885) sophiae, Helicarion sophiae L.A. Reeve, 1854 sophiae conica, Helicarion sophiae conica J.W. Brazier, 1889 sororcula, Trochochlamys sororcula H.A. Pilsbry & Y. Hirase, 1904 strangei, Helicarion strangei L. Pfeiffer, 1850 stylodon, Erepta stylodon L. Pfeiffer, 1842 subconvexa, Geotrochus conicoides subconvexa O.F. von Möllendorff subcrenulata, Helicarion subcrenulata H.A. Pilsbry, 1901 subcrenulata satsumana, Helicarion subcrenulata satsumana H.A. Pilsbry & Y. Hirase, 1908 subdetecta, Harmogenanina subdetecta J. Germain, 1921 subelimatus, Nipponochlamys subelimatus H.A. Pilsbry & Y. Hirase, 1904 submissus, Helicarion submissus T. Iredale, 1941 subrejectus, Helicarion subrejectus H.A. Pilsbry & Y. Hirase, 1908 subrejectus bella, Helicarion subrejectus bella H.A. Pilsbry, 1926 subrugatus, Nitor subrugatus L.A. Reeve, 1852 subtricolor, Geotrochus subtricolor (J.F. Mabille, 1887) superba, Helicarion superba J.C. Cox, 1871 suteri, Mathewsoconcha suteri E.R. Sykes, 1900 suturalis, Sesara suturalis N.H. Odhner, 1917 tagalensis, Lepidotrichia tagalensis (W.L.H. Dohrn, 1863) takahashii, Nipponochlamys takahashii J.T. Kuroda & T. Habe, 1969 takahashii, Trochochlamys fraterna takahashii J.T. Kuroda, 1963 tanegashimae, Yamatochlamys tanegashimae H.A. Pilsbry, 1901 tanzawaensis, Helicarion tanzawaensis E. Sorita & M. Kawana, 1983 tawale, Orpiella mouelti tawale Kae tenella, Orpiella tenella A. Garrett, 1872 tenellus, Helicarion tenellus A.A. Gould - syn of: Godwinia tenella (A.A. Gould) tenera, Helicarion tenera A. Adams, 1868 tentus, Westracystis tentus T. Iredale, 1939 - syn of: Westracystis lissus E.A. Smith, 1894 tenuisculpta, Nitor tenuisculpta R.B. Watson, 1873 teramachii, Helicarion teramachii J.T. Kuroda & H. Minato, 1976 testudo, Helicarion testudo H.B. Preston, 1913 tongkingensis, Helicarion tongkingensis O.F. von Möllendorff, 1901 - syn of: Megaustenia tongkingensis (O.F. von Möllendorff, 1901) unidentata, Nesonanina unidentata W.J. Clench, 1957 vaga, Yamatochlamys vaga H.A. Pilsbry & Y. Hirase, 1904 vaga adamsi, Yamatochlamys vaga adamsi J.T. Kuroda & T. Kawamoto, 1956 velutinella, Lepidotrichia velutinella (J.F. Quadras & O.F. von Möllendorff, 1892) venusta, Parakaliella venusta H.A. Pilsbry & Y. Hirase, 1909 vesta, Petalochlamys vesta L. Pfeiffer, 1865 vexillium, Mathewsoconcha vexillium H.B. Preston, 1913 virens, Helicarion virens L. Pfeiffer, 1849 vorticella, Ctenophila vorticella A. Adams, 1868 whiteheadi, Geotrochus whiteheadi (E.A. Smith, 1895) wiangariensis, Nitor wiangariensis I. Hyman, 2007 wolfi, Nesonanina wolfi C.R. Böttger, 1918 xenica, Helicarion xenica H.A. Pilsbry & Y. Hirase, 1903 zonatus, Geotrochus zonatus J.C. van Hasselt, 1823

Class: GASTROPODA --- Clade: HETEROBRANCHIA-PULMONATA-EUPULMONATA-STYLOMMATOPHORA-SIGMURETHRA-HELICARIONOIDEA ------ Family: HELICARIONIDAE Bourguignat, 1877 Habitat= Land, Taxonomic order Taxa=468, Genus=80, Subgenus=14, Species=311, Subspecies=20, Synonyms=42, Images=83

Genus: Bekkochlamys T. Habe, 1958

(4 SP, 4 IMG)

4322504291

Bekkochlamys dulcis H.A. Pilsbry, 1902 Japan, Kochi

4322504515

Bekkochlamys koshikijimana H.A. Pilsbry & Y. Hirase, 1904 Japan, Kagoshima

4322504691

Bekkochlamys micrograpta H.A. Pilsbry, 1900 Japan, Hyogo

5766000019

Bekkochlamys shikokuensis H.A. Pilsbry & Y. Hirase, 1903 Japan, Tokushima

synonyms: Macrochlamys perfragilis shikokuensis H.A. Pilsbry & Y. Hirase, 1903

Genus: Brevisentis I. Hyman, 2007

(1 SP, 0 IMG) 5766000027

Brevisentis atratus I. Hyman, 2007 New South Wales

Genus: Conibycus K.H.J. Thiele, 1928

(1 SP, 0 IMG)

5766000059

Conibycus dahli K.H.J. Thiele, 1928

No Georeferenced data available

Genus: Cucullarion J. Stanisic, 1998

(1 SP, 1 IMG)

5766000030

Cucullarion parkini J. Stanisic, 1998 Australia, Brisbane

Genus: Eungarion J. Stanisic, 1993

(1 SP, 0 IMG) 5766000031

Eungarion mcdonaldi J. Stanisic, 1993 Australia, Mackay

Genus: Howearion T. Iredale, 1944

(1 SP, 0 IMG) 4322506782

Howearion hilli (J.C. Cox, 1873) New South Wales

Genus: Hunsteinia T. Iredale, 1941

(5 SP, 4 IMG) 4322506774

Hunsteinia achilles (J.W. Brazier, 1886) Papua New Guinea

4322506776

Hunsteinia armiti (E.A. Smith, 1895) Papua New Guinea

4322506772

Hunsteinia flyensis (C. Hedley, 1894) Papua New Guinea 64,6 mm

4322506773

Hunsteinia foersteri (W. Kobelt, 1914) Papua New Guinea

5766000066

Hunsteinia hercules (C. Hedley, 1891)

No Georeferenced data available

Genus: Japanochlamys T. Habe, 1946

(1 SP, 1 IMG)

4322504251

Japanochlamys decens H.A. Pilsbry & Y. Hirase, 1904 Japan, Niigata

Genus: Kermarion E.A. Smith, 1873

(1 SP, 0 IMG) 5822000477

Kermarion kermadecensis (E.A. Smith, 1873) Kermadec Islands

Genus: Levidens I. Hyman, 2007

(1 SP, 0 IMG) 5766000028

Levidens ponderi I. Hyman, 2007 New South Wales

Genus: Lutilodix T. Iredale, 1945

(1 SP, 0 IMG) 5766000017

Lutilodix imitratrix E.R. Sykes, 1900 Norfolk Island

Genus: Nipponochlamys T. Habe, 1945

(8 SP, 4 IMG)

4322504403

Nipponochlamys hakusanus H.A. Pilsbry & Y. Hirase, 1907 Japan, Ishikawa

5766000023

Nipponochlamys hokkaidonis H.A. Pilsbry & Y. Hirase, 1905 Japan, Hokkaido

synonyms: Microcystina hokkaidonis H.A. Pilsbry & Y. Hirase, 1905

4322504475

Nipponochlamys izushichitojimana H.A. Pilsbry & Y. Hirase, 1904 Japan, Izu Islands

4322504595

Nipponochlamys lineatus H.A. Pilsbry & Y. Hirase, 1904 Japan

4322504755

Nipponochlamys obtusa C.A. Westerlund, 1883 Japan

4320504030

Nipponochlamys semisericata (H.A. Pilsbry, 1902) Wakayama, Japan

4322505059

Nipponochlamys subelimatus H.A. Pilsbry & Y. Hirase, 1904 Japan, Kagoshima

4322505091

Nipponochlamys takahashii J.T. Kuroda & T. Habe, 1969 Japan, Fukuoka

Genus: Ovachlamys T. Habe, 1946

(4 SP, 1 IMG) 4322504347

Ovachlamys fulgens G.P.L.K. Gude, 1900 Costa Rica

4322504355

Ovachlamys fulgida H.H. Godwin-Austen, 1907 Japan, Kagoshima

4322504491

Ovachlamys kandai M. Azuma, 1978 Japan

4322504523

Ovachlamys kotosyonis J.T. Kuroda & T. Kano, 1941 Japan

Genus: Panulena T. Iredale, 1945

(1 SP, 0 IMG) 4322504811

Panulena perrugosa T. Iredale, 1945 Norfolk Island, EXTINCT

Genus: Parakaliella T. Habe, 1946

(21 SP, 3 IMG) 4322504059

Parakaliella acutanguloides J.T. Kuroda & M. Azuma, 1982 Japan

4322504067

Parakaliella affinis H.A. Pilsbry & Y. Hirase, 1905 Japan, Hokkaido

4322504115

Parakaliella austeniana H.A. Pilsbry & Y. Hirase, 1901 Japan

5766000025

Parakaliella bimaris H.A. Pilsbry & Y. Hirase Japan, Okinawa

synonyms: Kaliella bimaris H.A. Pilsbry & Y. Hirase, 1904

4322504211

Parakaliella costata H.A. Pilsbry & Y. Hirase, 1904 Japan

4322504267

Parakaliella depressa O.F. von Möllendorff, 1883 Japan

4322504363

Parakaliella fusaniana H.A. Pilsbry & Y. Hirase, 1909 South Korea

4322504387

Parakaliella habei J.T. Kuroda, 1944 Japan, Wakayama

4322504427

Parakaliella hizenensis H.A. Pilsbry, 1902 Japan

4322504459

Parakaliella incerta A. Adams, 1868 Japan

4322204000

Parakaliella nahaensis (G.P.L.K. Gude, 1900) Okinawa

4322504763

Parakaliella obtusangula O.W.H. Reinhardt, 1883 Japan

4322504795

Parakaliella otakiana Y. Hirase, 1909 Japan

4322504803

Parakaliella pagoduloides G.P.L.K. Gude, 1900 Japan, Hyogo

4322504819

Parakaliella phyllophila A. Adams, 1868 Japan

4322504859

Parakaliella pupula A.A. Gould, 1859 Indo-Pacific

4322504867

Parakaliella pustulina O.W.H. Reinhardt, 1877 Japan

4322504939

Parakaliella ruida H.A. Pilsbry, 1901 Japan

4322504971

Parakaliella serica H.A. Pilsbry, 1927 Japan

4322504987

Parakaliella shikokuensis J.T. Kuroda & J.M. Abe, 1980 Japan

No Georeferenced data available

4322505155

Parakaliella venusta H.A. Pilsbry & Y. Hirase, 1909 Japan

Genus: Pseudosaphtia Winter, 2008

(1 SP, 0 IMG)

4322506778

Pseudosaphtia brunnea A.J. De Winter, 2008

No Georeferenced data available

Genus: Saphtia Winter, 2008

(2 SP, 0 IMG) 4322506780

Saphtia granulosa A.J. De Winter, 2008

No Georeferenced data available

4322506781

Saphtia lamtoensis A.J. De Winter, 2008

No Georeferenced data available

Genus: Sheaia I. Hyman, 2007

(1 SP, 0 IMG) 5766000029

Sheaia glenrockensis I. Hyman, 2007 New South Wales

Genus: Takemasaia M. Azuma & H. Minato 1976

(1 SP, 1 IMG)

4322504379

Takemasaia gudei H.A. Pilsbry, 1901 Hokkaido

Genus: Thularion J. Stanisic, 1993

(1 SP, 1 IMG) 4322506785

Thularion semoni E.C. Von Martens, 1894 Australia, Port Douglas

Genus: Trochochlamys T. Habe, 1946

(7 SP, 7 IMG)

4320504005

Trochochlamys crenulata G.P.L.K. Gude, 1900 Japan, Mitsumori

4322505203

Trochochlamys crenulata basistriata

H.A. Pilsbry & Y. Hirase, 1905 Japan

4322505211

Trochochlamys crenulata hotawana

H.A. Pilsbry & Y. Hirase, 1905 Japan

4322505427

Trochochlamys fraterna H.A. Pilsbry, 1900 Japan, Shimane

synonyms: Kaliella fraterna H.A. Pilsbry, 1900 4322505419

Trochochlamys fraterna takahashii J.T. Kuroda, 1963 Japan

5766000020

Trochochlamys humiliconus H.A. Pilsbry & Y. Hirase, 1904 Japan, Mie

synonyms: Kaliella humiliconus H.A. Pilsbry & Y. Hirase, 1904 4322504611

Trochochlamys lioconus H.A. Pilsbry & Y. Hirase, 1905 Japan, Yamagata

synonyms: Kaliella lioconus H.A. Pilsbry & Y. Hirase, 1905 4322504603

Trochochlamys lioconus goniozona

H.A. Pilsbry & Y. Hirase, 1905 Japan

4322504779

Trochochlamys okinoshimana H.A. Pilsbry & Y. Hirase, 1904 Japan, Kochi

synonyms: Kaliella okinoshimana H.A. Pilsbry & Y. Hirase, 1904 4322505955

Trochochlamys praealta H.A. Pilsbry, 1902 Japan, Niigata

synonyms: Kaliella praealta H.A. Pilsbry & Y. Hirase, 1903 Diastole praealta H.A. Pilsbry, 1902 4322505947

Trochochlamys praealta izushichtoensis

H.A. Pilsbry & Y. Hirase, 1903 Japan

4322505027

Trochochlamys sororcula H.A. Pilsbry & Y. Hirase, 1904 Japan, Tosa

synonyms: Kaliella sororcula H.A. Pilsbry & Y. Hirase, 1904

Genus: Westracystis T. Iredale, 1933

(2 SP, 2 IMG)

4322504331

Westracystis fredaslini G.A. Solem, 1982 Australia, Katherine 9,5 mm

4322504627

Westracystis lissus E.A. Smith, 1894 Australia, Weaber Ranges 6,7 mm

synonyms: Westracystis tentus T. Iredale, 1939

Genus: Yamatochlamys T. Habe, 1945

(2 SP, 2 IMG)

4322505099

Yamatochlamys tanegashimae H.A. Pilsbry, 1901 Japan, Kagoshima

synonyms: Microcystina tanegashimae H.A. Pilsbry, 1901 4322505147

Yamatochlamys vaga H.A. Pilsbry & Y. Hirase, 1904 Japan, Tokushima

synonyms: Microcystina vaga H.A. Pilsbry & Y. Hirase, 1904 4322505139

Yamatochlamys vaga adamsi

J.T. Kuroda & T. Kawamoto, 1956 Japan

SubFamily: HELICARIONINAE - Genus:

Bathia G.C. Robson, 1914

(1 SP, 1 IMG)

5766000018

Bathia madagascariensis G.C. Robson, 1914 Madagascar

SubFamily: HELICARIONINAE - Genus:

Caldwellia H. Adams, 1873

(2 SP, 1 IMG)

5766000010

Caldwellia imperfecta unknown Madagascar

5766000035

Caldwellia philyrina P.M.A. Morelet, 1851 Madagascar

SubFamily: HELICARIONINAE - Genus:

Ctenoglypta C.M.F. Ancey, 1904

(1 SP, 0 IMG)

5766000036

Ctenoglypta newtoni G. Nevill, 1871 Mauritius, EXTINCT

SubFamily: HELICARIONINAE - Genus:

Ctenophila C.M.F. Ancey, 1882 5766000013

Ctenophila caldwelli W.H. Benson, 1859 Mascarene Islands

(syn: Tachyphasis - 5 SP, 0 IMG)

5766000048

Ctenophila griffithsi K.C. Emberton, 1994 Madagascar

5822000518

Ctenophila milloti E. Fischer-Piette, F. Blanc, & F. Salvat, 1975 Madagascar

5766000014

Ctenophila setiliris W.H. Benson, 1851 Mascarene Islands

5766000037

Ctenophila vorticella A. Adams, 1868 Reunion

SubFamily: HELICARIONINAE - Genus:

Dancea A. Zilch, 1960

(1 SP, 1 IMG)

5766000012

Dancea rodriguezensis J.C.H. Crosse, 1873 Rodriguez Island, Mauritius

SubFamily: HELICARIONINAE - Genus:

Dendrotrochus H.A. Pilsbry, 1894 4322505315

Dendrotrochus cineraceus J.B. Hombron & C.H. Jacquinot, 1841 Solomon Islands

5766000128

Dendrotrochus cineraceus simboana (E.A. Smith, 1885) Solomon Islands

5792000067

Dendrotrochus cyrene (J.C.H. Crosse, 1865) Solomon Islands

(8 SP, 6 IMG)

synonyms: Helix cyrene J.C.H. Crosse, 1865 5766000065

Dendrotrochus filaris (M. Leschke, 1912)

No Georeferenced data available

5766000057

Dendrotrochus gracilior (B. Rensch, 1935)

No Georeferenced data available

4320954000

Dendrotrochus helicinoides J.B. Hombron & C.H. Jacquinot, 1841 Admiralty Islands

4322505323

Dendrotrochus helicinoides cleryi (C.A. Récluz, 1851)

Philippines, Palawan Island 6 mm

synonyms: Helix cleryi C.A. Récluz, 1851 4322505331

Dendrotrochus labillardierei E.A. Smith, 1884 Admiralty Islands

4322505339

Dendrotrochus layardi W.D. Hartman, 1889 Vanuatu 14,2 mm

4322505347

Dendrotrochus ponapensis H.B. Baker, 1941 Micronesia

SubFamily: HELICARIONINAE - Genus:

Dupontia H.H. Godwin-Austen, 1908

(7 SP, 0 IMG)

5766000038

Dupontia levensonia J. Gerlach, 2003 Seychelles

5766000039

Dupontia levis H.H. Godwin-Austen, 1908 Mauritius

5766000052

Dupontia maillardi (G.P. Deshayes, 1863) Reunion

5766000040

Dupontia nitella P.M.A. Morelet, 1868 Mauritius

5766000015

Dupontia perlucida A. Adams, 1867 Mascarene Islands

5766000041

Dupontia poweri A. Adams, 1868 Mauritius

5766000042

Dupontia proletaria P.M.A. Morelet, 1860 Mauritius, EXTINCT

SubFamily: HELICARIONINAE - Genus:

Epiglypta H.A. Pilsbry, 1893 4322505179

Epiglypta howinsulae J.C. Cox, 1873 Lord Howe Island, Australia

(1 SP, 0 IMG)

SubFamily: HELICARIONINAE - Genus:

Erepta J.A. Albers, 1850

(syn: Microstylodonta - 3 SP, 0 IMG)

5766000122

Erepta nevilli A. Adams, 1867 Mauritius, EXTINCT

5766000016

Erepta odontina P.M.A. Morelet, 1851 Mascarene Islands

5766000043

Erepta stylodon L. Pfeiffer, 1842 Mauritius

SubFamily: HELICARIONINAE - Genus:

Geotrochus J.C. Van Hasselt, 1823 5766000119

Geotrochus affinis (E.A. Smith, 1894) Indonesia, Borneo

5766000090

Geotrochus alexis (E.A. Smith, 1895) Indonesia, Borneo

5766000091

Geotrochus bongaoensis (E.A. Smith, 1894) Indonesia, Borneo

4322505355

Geotrochus conicoides W. Metcalfe, 1851 Indonesia, Borneo

5766000092

Geotrochus conicoides parva

(E.A. Smith, 1887) Indonesia, Borneo

(syn: Chiroktisma - 20 SP, 3 IMG)

5766000005

Geotrochus conicoides subconvexa

O.F. von Möllendorff Philippines, Palawan Island 15,8 mm

5766000062

Geotrochus conus (R.A. Philippi, 1841)

No Georeferenced data available

5766000093

Geotrochus everetti (H.H. Godwin-Austen, 1891) Indonesia, Borneo

5766000094

Geotrochus heraclea (E.A. Smith, 1895) Indonesia, Borneo

5792000092

Geotrochus isabellensis (S.M. Souverbie, 1863) Solomon Islands 17 mm

synonyms: Helix isabellensis S.M. Souverbie, 1863 5766000095

Geotrochus kinabaluensis (E.A. Smith, 1895) Indonesia, Borneo

4322505363

Geotrochus labuanensis L. Pfeiffer, 1863 Indonesia, Borneo

5766000096

Geotrochus meristotrochus J.J. Vermeulen, M. Schilthuizen, & T.S. Liew MS Indonesia, Borneo

5766000097

Geotrochus obscura J.J. Vermeulen, M. Schilthuizen, & T.S. Liew MS Indonesia, Borneo

5766000098

Geotrochus pallida (E.A. Smith, 1895) Indonesia, Borneo

5766000099

Geotrochus paraguensis (E.A. Smith, 1893) Indonesia, Borneo

5766000100

Geotrochus rhysa S. Tillier & P. Bouchet, 1988 Indonesia, Borneo

5766000053

Geotrochus rimatus J.J. Vermeulen, 1997

No Georeferenced data available

5822000468

Geotrochus saigonensis (J.C.H. Crosse, 1867) South Vietnam

synonyms: Trochomorpha saigonensis J.C.H. Crosse, 1867 5822000471

Geotrochus subtricolor (J.F. Mabille, 1887) North Vietnam

5766000101

Geotrochus whiteheadi (E.A. Smith, 1895) Indonesia, Borneo

4322505371

Geotrochus zonatus J.C. van Hasselt, 1823 Indonesia, West Java 15 mm

synonyms: Helix conus R.A. Philippi, 1841 SubFamily: HELICARIONINAE - Genus:

Harmogenanina J. Germain, 1919

(syn: Cycliscus, Rotula - 5 SP, 1 IMG)

5766000044

Harmogenanina argentea L.A. Reeve, 1852 Reunion

5766000011

Harmogenanina detecta (A.E.J. Férussac, 1827) Rodrigues Island

5766000045

Harmogenanina implicata G. Nevill, 1870 Mauritius

5766000123

Harmogenanina linophora P.M.A. Morelet, 1860 Mauritius, EXTINCT

5766000124

Harmogenanina subdetecta J. Germain, 1921 Reunion, EXTINCT

SubFamily: HELICARIONINAE - Genus: Vercularion - 102 SP, 7 IMG)

Helicarion A.E.J. Férussac, 1821 4322504043

Helicarion aberrans T. Iredale, 1944 Lord Howe Island, Australia

(syn: Desidarion, Fastosarion, Helixarion, Helicarium, Laconia, Luinarion, Mysticarion, Parmavitrina, Peloparion,

4322504051

Helicarion acutangula A. Adams, 1868 Japan

4322504075

Helicarion akibumii J.T. Kuroda, 1963 Japan

5766000008

Helicarion albellus E.C. Von Martens, 1867 Java, Indonesia 8 mm

4322504083

Helicarion alma T. Iredale, 1944 Lord Howe Island, Australia

4322504091

Helicarion amiculus T. Iredale, 1945 Norfolk Island

4322504099

Helicarion ampla O.W.H. Reinhardt, 1883 Japan

4322506763

Helicarion aquavitae P.H. Scott, 1995 Mackay, Australia 8 mm

4322504107

Helicarion aquila J.C. Cox, 1868 New South Wales

4322504123

Helicarion awaensis H.A. Pilsbry, 1902 Japan

4322504131

Helicarion azumai T. Habe, 1948 Japan

4322504139

Helicarion belli T. Iredale, 1944 Lord Howe Island, Australia

4322504147

Helicarion borealis H.A. Pilsbry, 1901 Indo-Pacific

4322504155

Helicarion brazieri J.C. Cox, 1873 Australia, Palm Island

4322504163

Helicarion bullaceus N.H. Odhner, 1917 Queensland

4322504171

Helicarion castanea L. Pfeiffer, 1853 Great Australian Bight

4322504179

Helicarion catletti J.W. Brazier, 1872 Lord Howe Island, Australia

No Georeferenced data available

4322504187

Helicarion ceratodes G.P.L.K. Gude, 1900 Japan

4322504195

Helicarion charis T. Iredale, 1944 Lord Howe Island, Australia

4322504203

Helicarion circumdata H.A. Pilsbry, 1902 Japan, Tokyo

4322504219

Helicarion crassiaxis J.T. Kuroda, 1969 Japan

4322504227

Helicarion crenata J.T. Kuroda & M. Azuma, 1982 Japan

4322504235

Helicarion cuvieri A.E.J. Férussac, 1821 Tasmania

4322504243

Helicarion danjoensis H. Minato, 1974 Japan, Nagasaki

4322504259

Helicarion delecta T. Iredale, 1944 Lord Howe Island, Australia

4322504275

Helicarion depressus G.A. von Jacobi, 1898 Japan

4322504283

Helicarion dispositus T. Iredale, 1941 Australia, Gloucester

4322504299

Helicarion dulcissima T. Iredale, 1944 Lord Howe Island, Australia

4322504307

Helicarion etheridgei J.W. Brazier, 1889 Lord Howe Island, Australia

4322504315

Helicarion flavescens T. Iredale, 1944 Lord Howe Island, Australia

4322504323

Helicarion fraternus T. Iredale, 1945 Norfolk Island

4322504339

Helicarion freycineti A.E.J. Férussac, 1821 New South Wales

4322504371

Helicarion grata T. Iredale, 1944 Lord Howe Island, Australia

4322504395

Helicarion hakonensis H.A. Pilsbry & Y. Hirase, 1905 Japan, Kanagawa

4322504411

Helicarion helenae H.H. Godwin-Austen, 1883 Australia, Sydney

5766000032

Helicarion helenkingae P.H. Scott, 1995 Australia, Townsville

4322504419

Helicarion hiraseana H.A. Pilsbry, 1901 Congo

4322504435

Helicarion hyaleus F.S. Bock, 1881 Indonesia

4322504443

Helicarion imitatrix E.R. Sykes, 1900 Norfolk Island

4333454396

Helicarion imperator (A.A. Gould, 1859) China, Guangxi 31 mm

synonyms: Vitrina imperator A.A. Gould, 1859 Helicarion maleficus P. Dautzenberg, 1904

Megaustenia imperator A.A. Gould, 1859

5792000164

Helicarion imperator brunneus

(O.F. von Möllendorff, 1901) North Vietnam

5766000102

Helicarion infans (L. Pfeiffer, 1854) Indonesia, Borneo

4322504467

Helicarion insculpta L. Pfeiffer, 1846 Norfolk Island

5766000034

Helicarion iridis I. Hyman, 2007 Kyogle, Australia

4322504483

Helicarion kagaensis H.A. Pilsbry & Y. Hirase, 1902 Japan, Ishikawa

4322504499

Helicarion kiiensis M. Azuma, 1960 Japan

4322504507

Helicarion kimioi M. Azuma, 1960 Japan

4322504531

Helicarion koyamai J.T. Kuroda, 1953 Japan

4322504539

Helicarion koyanoi T. Habe, 1953 Japan

4322504547

Helicarion kreffti J.C. Cox, 1864 Queensland

4322504555

Helicarion kurodai T. Habe, 1957 Japan

4322504563

Helicarion kuroshimana Y. Hirase, 1908 Japan

4322504571

Helicarion lampra H.A. Pilsbry & Y. Hirase, 1904 Japan, Hyogo

synonyms: Microcystina lampra H.A. Pilsbry & Y. Hirase, 1904 4322504579

Helicarion leopardina T. Iredale, 1941 New South Wales

4322504587

Helicarion leucospira L. Pfeiffer, 1857 Australia, Lismore

4322504619

Helicarion lioderma H.A. Pilsbry, 1901 Japan

4322504635

Helicarion longa H.A. Pilsbry & Y. Hirase, 1905 Japan

4322504643

Helicarion longissima H.A. Pilsbry & Y. Hirase, 1909 Japan

4322504651

Helicarion masakii J.T. Kuroda, 1960 Japan

4322504667

Helicarion mastersi J.C. Cox, 1868 Australia, Kiama 13 mm

4322504659

Helicarion mastersi callidus

T. Iredale, 1941 New South Wales

4322504675

Helicarion mathewsi H.B. Preston, 1913 Norfolk Island

4322504683

Helicarion megastoma J.C. Cox, 1868 New South Wales

4322504699

Helicarion miranda T. Iredale, 1944 Lord Howe Island, Australia

4322504707

Helicarion monticula J.T. Kuroda & J.M. Abe, 1980 Japan, Tokushima

synonyms: Helicarion monticola J.T. Kuroda & J.M. Abe, 1980

4322504723

Helicarion mutsuensis Y. Hirase, 1908 Japan

4322504731

Helicarion nachikoae J.T. Kuroda, 1953 Japan

4322504739

Helicarion nigra J.R.C. Quoy & J.P. Gaimard, 1832 Victoria

4322504747

Helicarion nikkoensis E. Sorita & M. Kawana, 1983 Japan

5766000002

Helicarion novoguineensis E.A. Smith, 1887 Papua New Guinea

4322504771

Helicarion ogasawarana H.A. Pilsbry, 1902 Japan

4322504787

Helicarion osimakiheijii J.T. Kuroda, 1953 Japan

4322504827

Helicarion planilabris J.C. Cox, 1866 New South Wales

5766000078

Helicarion planospirus (L. Pfeiffer, 1853)

No Georeferenced data available

4322504835

Helicarion porrectus T. Iredale, 1941 Australia, Dungog

4322504843

Helicarion prensa T. Iredale, 1944 Lord Howe Island, Australia

4322504851

Helicarion princeps T. Iredale, 1944 Lord Howe Island, Australia

4322504875

Helicarion quelpartensis H.A. Pilsbry & Y. Hirase, 1902 South Korea

4322504883

Helicarion quintalae J.C. Cox, 1870 Norfolk Island

5766000080

Helicarion radiatulus (O.F. von Möllendorff, 1897)

No Georeferenced data available

4322504891

Helicarion rapida J.T. Kuroda & M. Miyanaga, 1939 Japan

4322504899

Helicarion retinaculum H.B. Preston, 1913 Norfolk Island

4322504907

Helicarion robusta A.A. Gould, 1846 New South Wales

4322504915

Helicarion rosacea T. Iredale, 1944 Lord Howe Island, Australia

4322504923

Helicarion rubicundus Dartnall & J.A. Kershaw, 1978 Tasmania

4322504931

Helicarion rugosa H.C. Fulton, 1910 Indonesia, Borneo

4322504947

Helicarion satsumana Y. Hirase, 1908 Japan

5766000103

Helicarion semicalcareus M.M. Schepman, 1896 Indonesia, Borneo

4322504955

Helicarion septentrionalis G.A. von Jacobi, 1898 Japan

4322504963

Helicarion serenus H.A. Pilsbry & Y. Hirase, 1908 Japan, Yamagata

4322504995

Helicarion shinanoensis H.A. Pilsbry & Y. Hirase, 1904 Japan

4322505003

Helicarion shiroyamaensis J.T. Kuroda & T. Habe, 1954 Japan

4322505019

Helicarion sophiae L.A. Reeve, 1854 Lord Howe Island, Australia

4322505011

Helicarion sophiae conica

J.W. Brazier, 1889 Lord Howe Island, Australia

4322505035

Helicarion strangei L. Pfeiffer, 1850 Queensland

4322505051

Helicarion subcrenulata H.A. Pilsbry, 1901 Japan, Kagoshima

4322505043

Helicarion subcrenulata satsumana

H.A. Pilsbry & Y. Hirase, 1908 Japan

4322505067

Helicarion submissus T. Iredale, 1941 New South Wales

4322505075

Helicarion subrejectus H.A. Pilsbry & Y. Hirase, 1908 Japan, Nagasaki

4322505259

Helicarion subrejectus bella

H.A. Pilsbry, 1926 South Korea

4322505083

Helicarion superba J.C. Cox, 1871 Guangxi, China 25 mm

4322505107

Helicarion tanzawaensis E. Sorita & M. Kawana, 1983 Japan

4322505115

Helicarion tenera A. Adams, 1868 Indo-Pacific

4322505123

Helicarion teramachii J.T. Kuroda & H. Minato, 1976 Japan, Fukuoka

4322505131

Helicarion testudo H.B. Preston, 1913 Norfolk Island 4,6 mm

No Georeferenced data available

4322505163

Helicarion virens L. Pfeiffer, 1849 Australia, Clarence River

4322505171

Helicarion xenica H.A. Pilsbry & Y. Hirase, 1903 Japan

SubFamily: HELICARIONINAE - Genus:

Hemiglypta O.F. von Möllendorff, 1893

(9 SP, 7 IMG)

4320954005

Hemiglypta blainvilleana H.C. Lea Philippines, Catanduanes 50 mm

4322506787

Hemiglypta cuvierana (H.C. Lea, 1893) Philippines, Luzon Island

4322505379

Hemiglypta globosa C.G. Semper, 1870 Philippines, Jabonga

4322505387

Hemiglypta iloilana M. Smith, 1932 Philippines

5766000070

Hemiglypta microglypta O.F. von Möllendorff, 1893

No Georeferenced data available

4320954015

Hemiglypta panayensis W.J. Broderip, 1842 Philippines, Mindoro Island 43 mm

4322506755

Hemiglypta panayensis connectens

O.F. von Möllendorff, 1898 Philippines, Surigao 90,3 mm

4322506771

Hemiglypta semiglobosa (L. Pfeiffer) Philippines, Romblon Island 42,1 mm

4322506751

Hemiglypta semigranosa G.B. I Sowerby, 1832 Philippines, Palawan Island 50 mm

4320954020

Hemiglypta semperi O.F. von Möllendorff Philippines, Palawan Island 39 mm

SubFamily: HELICARIONINAE - Genus:

Hemiglyptopsis K.H.J. Thiele, 1931

(1 SP, 0 IMG)

4322505395

Hemiglyptopsis fouillioyi E.J. Le Guillou, 1845 Papua New Guinea

SubFamily: HELICARIONINAE - Genus:

Inozonites G.J. Pfeffer, 1883

(1 SP, 0 IMG)

5766000075

Inozonites orthostoma (L. Pfeiffer, 1845)

No Georeferenced data available

SubFamily: HELICARIONINAE - Genus:

Lepidotrichia P. Bartsch, 1942

(7 SP, 1 IMG)

5766000068

Lepidotrichia kobelti (O.F. von Möllendorff, 1890)

No Georeferenced data available

5766000056

Lepidotrichia pandana (P. Bartsch, 1942)

No Georeferenced data available

5766000079

Lepidotrichia pruinosa (O.F. von Möllendorff, 1890)

No Georeferenced data available

4322505507

Lepidotrichia purpurescens O.F. von Möllendorff Philippines

4322506792

Lepidotrichia semisculpta O.F. von Möllendorff Philippines 24 mm

5766000085

Lepidotrichia tagalensis (W.L.H. Dohrn, 1863)

No Georeferenced data available

5766000088

Lepidotrichia velutinella (J.F. Quadras, & O.F. von Möllendorff, 1892)

SubFamily: HELICARIONINAE - Subgenus:

No Georeferenced data available

Lepidotrichia (Hemitrichiella) A. Zilch, 1956

4321554000

Hemitrichiella baleriana P. Bartsch, 1939 Philippines 23 mm

4322506767

Hemitrichiella guamata G.B. Sowerby Philippines, Batanes Island 36,7 mm

(syn: Hemitrichia - 4 SP, 5 IMG)

4322505499

Hemitrichiella luteofasciata H.C. Lea, 1840 Philippines, Luzon Island 27,4 mm

4322505491

Hemitrichiella luteofasciata libmanana P. Bartsch, 1942

Philippines, Catanduanes 30 mm

4321554005

Hemitrichiella setigera G.B. Sowerby, 1898 Philippines, Palawan Island 22,1 mm

SubFamily: HELICARIONINAE - Genus:

Mathewsoconcha H.B. Preston, 1913 4322505515

Mathewsoconcha albocincta H.B. Preston, 1913 Norfolk Island

4322505523

Mathewsoconcha belli H.B. Preston, 1913 Norfolk Island

4322505531

Mathewsoconcha compacta H.B. Preston, 1913 Norfolk Island

4322505539

Mathewsoconcha elevata H.B. Preston, 1913 Norfolk Island

4322505547

Mathewsoconcha microstriata H.B. Preston, 1913 Norfolk Island

(8 SP, 1 IMG)

4322505555

Mathewsoconcha phillipii J.E. Gray, 1834 Norfolk Island

4322505563

Mathewsoconcha suteri E.R. Sykes, 1900 Norfolk Island

4322505571

Mathewsoconcha vexillium H.B. Preston, 1913 Norfolk Island

SubFamily: HELICARIONINAE - Genus:

Nesonanina C.R. Böttger, 1916

(4 SP, 2 IMG)

4322505579

Nesonanina malleata B. Rensch Papua New Guinea

4322505587

Nesonanina novaehiberniae J.R.C. Quoy & J.P. Gaimard, 1832 New Ireland

4322505595

Nesonanina unidentata W.J. Clench, 1957 Papua New Guinea

4322505603

Nesonanina wolfi C.R. Böttger, 1918 New Ireland

SubFamily: HELICARIONINAE - Genus:

Nitor G.P.L.K. Gude, 1911

(syn: Modonitor, Pravinitor, Thalassia - 7 SP, 0 IMG)

4322505611

Nitor coriacea A. Manzoni, 1868 Europe

No Georeferenced data available

4322505619

Nitor dagueneti L. de Folin, 1873 Europe

No Georeferenced data available

4322505627

Nitor medioximus T. Iredale, 1941 Australia, Port Macquarie

4322505635

Nitor pudibunda J.C. Cox, 1868 Australia, Richmond

4322505643

Nitor subrugatus L.A. Reeve, 1852 Australia, Clarence River

4322505651

Nitor tenuisculpta R.B. Watson, 1873 Europe

No Georeferenced data available

5766000033

Nitor wiangariensis I. Hyman, 2007 Australia, Brisbane

SubFamily: HELICARIONINAE - Genus:

Orpiella J.E. Gray, 1855 4322505659

Orpiella compluviata (J.C. Cox, 1871) Solomons Islands, Guadalcanal Island

4322506793

Orpiella cookensis A.A. Gould Fiji Islands 25 mm

(syn: Eurypus, Fretum - 10 SP, 5 IMG)

4322506794

Orpiella fragillima A.A. Gould Fiji Islands 13 mm

4322505667

Orpiella godeffroyana A. Garrett, 1872 Fiji Islands

5766000001

Orpiella mouelti Kae

No Georeferenced data available

4322506795

Orpiella mouelti tawale Kae

Fiji Islands 29 mm

5766000073

Orpiella nouleti (E.J. Le Guillou, 1842)

No Georeferenced data available

5766000077

Orpiella pfeifferi (R.A. Philippi, 1845)

No Georeferenced data available

4322505675

Orpiella scorpio A.A. Gould Fiji Islands

5766000082

Orpiella solidiuscula (E.A. Smith, 1885)

No Georeferenced data available

5766000024

Orpiella tenella A. Garrett, 1872 Fiji Islands

synonyms: Nanina tenella A. Garrett, 1872 SubFamily: HELICARIONINAE - Subgenus:

Orpiella (Irenella) G.P.L.K. Gude, 1913

(syn: Aulacopus - 2 SP, 2 IMG)

5766000021

Irenella hoyti A. Garrett, 1872 Fiji Islands

synonyms: Nanina hoyti A. Garrett, 1872 5766000022

Irenella otareae A. Garrett Fiji Islands

SubFamily: HELICARIONINAE - Subgenus:

Orpiella (Owaraha) F.C. Baker, 1941

5766000009

Owaraha nissani R.K. Dell, 1955 New Zealand

SubFamily: HELICARIONINAE - Genus:

Pachystyla O.A.L. Mörch, 1852

(2 SP, 1 IMG)

4322505827

Pachystyla inversicolor A.E.J. Férussac, 1821 Mauritius

5766000127

Pachystyla inversicolor mauritianella (P.M.A. Morelet) Mauritius 16 mm

5766000125

Pachystyla rufozonata A. Adams, 1867 Mauritius, EXTINCT

SubFamily: HELICARIONINAE - Genus:

Parmacochlea E.A. Smith, 1884

(2 SP, 0 IMG)

(1 SP, 1 IMG)

4322505187

Parmacochlea fischeri E.A. Smith, 1884 Queensland

4322505195

Parmacochlea smithi H.R. Simroth, 1898 Australia

SubFamily: HELICARIONINAE - Genus:

No Georeferenced data available

Petalochlamys H.H. Godwin-Austen, 1907 4320504000

Petalochlamys cerasina (H.A. Pilsbry, 1902) Japan, Mitsumori

4320504010

Petalochlamys doenitzii (O.W.H. Reinhardt, 1877) Japan, Minamiamabe

4322505227

Petalochlamys formosana P.B. Schmacker & O. Böttger, 1891 Taiwan

synonyms: Petalochlamys formosana H.A. Pilsbry & Y. Hirase, 1908 4320504015

Petalochlamys labilis (A.A. Gould, 1859) Japan, Mitsumori

4322505235

Petalochlamys nitidus H.A. Pilsbry & Y. Hirase, 1905 Taiwan

4322505243

Petalochlamys par P.B. Schmacker & O. Böttger, 1891 Japan

(9 SP, 5 IMG)

4320504020

Petalochlamys perfragilis (H.A. Pilsbry, 1901) Japan, Kagoshima

4320504025

Petalochlamys perfragilis sakui

H.A. Pilsbry & Y. Hirase, 1908 Japan, Tanegashima Island

4322505251

Petalochlamys rejecta L. Pfeiffer, 1865 China, Zhejiang

4322505267

Petalochlamys vesta L. Pfeiffer, 1865 Taiwan

SubFamily: HELICARIONINAE - Genus:

Plegma G.P.L.K. Gude, 1911

(1 SP, 1 IMG)

5766000126

Plegma caelatura (A.E.J. Férussac, 1821) French Guiana, Cayenne

SubFamily: HELICARIONINAE - Genus:

Pseudaustenia T.D.A. Cockerell, 1891

(1 SP, 1 IMG)

5766000121

Pseudaustenia atra H.H. Godwin-Austen India, Western Ghats

SubFamily: HELICARIONINAE - Genus:

Rahula H.H. Godwin-Austen, 1907

(2 SP, 0 IMG)

5766000046

Rahula moolenbeeki W.J.M. Maassen, 2000

No Georeferenced data available

5766000120

Rahula raricostulata (E.A. Smith, 1894) Indonesia, Borneo

SubFamily: HELICARIONINAE - Genus:

Roybellia H.B. Preston, 1913

(2 SP, 0 IMG)

4322505683

Roybellia depressa H.B. Preston, 1913 Norfolk Island

4322505691

Roybellia platysoma E.R. Sykes, 1900 Norfolk Island

SubFamily: HELICARIONINAE - Genus:

Sesara J.A. Albers, 1880 5792000161

Sesara annamitica (O.F. von Möllendorff, 1900) Vietnam, Da Nang

5792000163

Sesara bouyei (J.C.H. Crosse & P.H. Fischer, 1863) South Vietnam

synonyms: Ariophanta bouyei J.C.H. Crosse & P.H. Fischer, 1863 5766000049

Sesara diplodon (W.H. Benson, 1859) Bangladesh

4322505283

Sesara infredens A.A. Gould Myanmar

(syn: Malandena - 6 SP, 0 IMG)

5792000165

Sesara infrendens (A.A. Gould, 1844) South Vietnam

synonyms: Helix infrendens A.A. Gould, 1844 4322505291

Sesara suturalis N.H. Odhner, 1917 Queensland

SubFamily: HELICARIONINAE - Genus:

Sivella W.T. Blanford, 1863

(4 SP, 0 IMG)

5792000160

Sivella albofilosa (A.R.J.B. Bavay & P. Dautzenberg, 1908) North Vietnam

5792000166

Sivella latior (A.R.J.B. Bavay & P. Dautzenberg, 1908) Vietnam, Nat Son

synonyms: Trochomorpha latior A.R.J.B. Bavay & P. Dautzenberg, 1908 5792000168

Sivella montana (O.F. von Möllendorff, 1901) North Vietnam

synonyms: Trochomorpha montana O.F. von Möllendorff, 1901 5792000171

Sivella paviei (L.J. Morlet, 1884) North Vietnam

synonyms: Trochomorpha tonkinorum L.J. Morlet, 1884 SubFamily: HELICARIONINAE - Genus:

Wilhelminaia H.B. Preston, 1913

(1 SP, 0 IMG)

4322505275

Wilhelminaia mathildae H.B. Preston, 1913 Brasil, Espirito Santo

SubFamily: PAPUARIONINAE - Genus:

Laocaia A.A. Kuzminykh, 1999

(2 SP, 0 IMG)

5766000006

Laocaia attenuata A.A. Kuzminykh, 1999 Vietnam

5792000170

Laocaia obesa A.A. Kuzminykh, 1999 North Vietnam

Family: HELICARIONIDAE

red squares show where each species has been georeferenced each square has a size of 2 degrees an interactive version of this image can be accessed at http://www.bagniliggia.it/WMSD/FamilyMap/HELICARIONIDAEmap.htm please note that this is not the complete coverage of this family the Data Base is not exaustive and only one specimen per species has been georeferenced this map is only the graphic summary of the WMSDB Data Base up to now the total number of georeferenced species in this map is: 301 from a total of 331

Author: Claudio Galli - [email protected]